Haminoea miocaenica

Zenodo (CERN European Organization for Nuclear Research)

Abstract

Haminoea miocaenica (Berger, 1949) Figs 48A–C Bulla hydatis L.— Friedberg 1928: 547, pl. 36, fig. 5 [non Haminoea hydatis (Linnaeus, 1758)]. * Haminea (Haminea) hydatis miocaenica n. subsp. — Berger 1949: 82. Haminea hydatis miocaenica n. subsp. — Berger 1953: 87, pl. 16, figs 5–7. H [aminea]. hydatis miocaenica Berg.— Sieber 1958: 163. Haminoea hidatis [sic] (Linné, 1758)— Zelinskaya et al. 1968: 234, pl. 52, figs 25–26 [non Haminoea hydatis (Linnaeus, 1758)]. Haminea hydatis miocaenica Berger— Bałuk 1970: 119, pl. 14, fig. 4. Haminoaea [sic] hydatis L.—Rado & Muţui 1970: 148, pl. 6, figs 17–19 [non Haminoea hydatis (Linnaeus, 1758)]. Haminoea cf. hydatis miocaenica Berger, 1953 — Landau et al. 2013, 327, pl. 52, fig. 17. Type material. Holotype, NHMW-ZOO-MO-75000-E-3290, SL: 3.4 mm, MD: 2.4 mm, Bad Vöslau (Austria), illustrated in Berger (1953: pl. 16, fig. 7), Figs 48A 1 –A 3. Paratype, NHMW-ZOO-MO-75000-E-36704. SL: 3.4 mm, MD: 2.3 mm, Guntersdorf (Austria), Figs 48B 1 –B 3. Illustrated material. NHMW 1997z01781213, SL: 6.9 mm, MD: 4.6 mm, Bad Vöslau (Austria), Figs 48C 1 – C 3. Revised description. Shell medium sized, delicate, glossy, very broad, ovate-subcylindrical (SL/MD = 1.5). Only last whorl visible; periphery placed just above mid-whorl (SL/ PH = 1.1). Broad, rounded apex (MD/ApD = 1.4), with moderately oblique apical plane (AP = 12°), moderately wide, shallow umbilicus. Sculpture of weak growth lines. Aperture relatively wide in anterior third (MD/AW = 4.1), greatly widening anteriorly (MD/MAW = 2.0), resulting in low expansion rate (ER = 2.1). Outer lip thin, only slightly rising above apex, weakly convex at periphery, rounded abapically. Columella short, strongly excavated in lower half. Columellar lip forming narrow rim, delimiting narrow umbilical chink. Discussion. We follow Berger (1949, 1953) and Landau et al. (2013) and separate the Middle Miocene specimens from Haminoea hydatis (Linnaeus, 1758), which is larger, has a broader, fully adherent columellar callus and an outer lip usually raised above the apex. However, more Miocene specimens will be needed to decide if these features really justify separation from the Pliocene to extant species. Haminoea hydatis var. hydatina (Dod.), from the Tortonian of Montegibbio (Italy) as illustrated by Sacco (1897: pl. 4, fig 6), was listed by Landau et al. (2013) as synonym of Haminoea miocaenica (Berger, 1949). This name would have priority over Haminoea miocaenica, but the fragmentary Italian specimen does not even allow a clear generic identification. Friedberg (1928) listed Bulla inflata Eichwald, 1830, from the Middle Miocene of Ukraine, as synonym of this species (as Bulla hydatis). We have not seen the type specimens of ‘ Bulla ’ inflata and its status remains unclear. The illustration in Eichwald (1852: pl. 11, fig 14) shows an ovate-conical shell with its periphery distinctly below mid-whorl, with narrow adapical aperture and deeply excavated columella, which is quite unlike Haminoea miocaenica (Berger, 1949). Therefore, we tentatively remove it from the chresonymy but note that this name would also have priority over Haminoea miocaenica. Paleoenvironment. The occurrence in the Baden Formation at Bad Vöslau (Austria) suggests middle to outer neritic environments with up to 210 m water depth (Kranner et al. 2021), whereas the occurrences at Steinebrunn (Austria) and Seyithasan (Turkey) point to shallow water environments (own data M.H., B.L.). Distribution in Central Paratethys. Badenian (Middle Miocene): Polish-Ukrainian Fore-Carpathian Basin Wieliczka (Poland), Faszczówka, Staryi Pochaiv, Shushkivtsi, Zhukivtsi (Ukraine) (Friedberg 1938); Voronyaky Hills: Oles’ko (Ukraine) (Friedberg 1938); North Alpine-Carpathian Foreland Basin: Hrušovany nad Jevišovkou (Czech Republic), Guntersdorf (Austria) (Berger 1953); Vienna Basin: Bad Vöslau, Steinebrunn, Gaaden (Austria) (Berger 1953). Dacian Basin: Islaz (Romania) (Rado & Muţui 1970). Proto-Mediterranean Sea. Serravallian (Middle Miocene): Karaman Basin: Seyithasan (Turkey) (Landau et al. 2013).

Authors 3

1. Mathias Harzhauser

   Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655

2. Bernard Landau Aachen

   University of Lisbon · Instituto Dom Luiz · Naturalis Biodiversity Center

   Affiliation as printed

   Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494

3. Manuel António E. Malaquias

   University of Bergen · Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655 & Department of Natural History, University Museum of Bergen, University of Bergen, PB 7800, 5020 - Bergen, Norway manuel. malaquias @ uib. no; http: // orcid. org / 0000 - 0002 - 9668 - 945 X & Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655