Hermania scabra

Abstract

Hermania scabra (Müller, 1784) Figs 54A–C * Bulla scabra — Müller 1784: 90, figs 1–3. Bullaea scabra Müll.— Wood 1848: 181, pl. 21, fig. 12. Hermania scabra (Müll.) — Cossmann 1895: 128, pl. 7, figs 1–2. Philine (Hermania) scabra var. pliocenica Sacc.— Sacco 1897: 54, pl. 4, fig. 42. Philine (Hermania) scabra (Müll.) —Boettger 1907: 207. Philine scabra pliocaenica [sic] Sacco, 1897 — Csepreghy-Meznerics 1952: 229, pl. 14, figs 13, 14. Philine (Hermania) scabra (Müller) — Berger 1953: 115, pl. 18, figs 83–86. P [hiline]. (H [ermania].) scabra (Müll.) — Sieber 1958: 164. Philine scabra Müller, 1776 [sic]— Glibert, 1960: 25, pl. 4, fig. 29. Philine scabra pliocaenica [sic] Sacco— Strausz 1962: 156, pl. 75, figs 12, 13. Philine scabra pliocaenica [sic] Sacco, 1897 — Strausz 1966: 483, pl. 75, figs 12, 13. Philine (Hermania) scabra (Müller, 1776 [sic])— Janssen 1984: 374, pl. 19, fig. 12. Philine scabra (Müller, 1776 [sic])— Landau et al. 2011: 42, pl. 23, fig. 11. Philine scabra (Müller, 1784)— Cossignani & Ardovini 2011: 38, 378, unnumbered fig. Philine scabra (Müller, 1784)— Gofas et al. 2011: 411, unnumbered fig. bottom. Philine scabra (Müller, 1784) — Chirli 2013: 34, pl. 9, figs 6–11. Philine scabra (Müller, 1784) — Landau et al. 2013: 330, pl. 77, fig. 4. Philine scabra (Müller, 1784) — Negri & Corselli 2016: 84, figs 19k–n. Hermania scabra (Müller, 1784) — Landau et al. 2023: 398: pl. 47, fig. 1 [cum syn.]. Type material. We are not aware of the whereabouts of the type material. Illustrated material. NHMW 2010/0004/1648, SL: 3.7 mm, MD: 2.3 mm, Bad Vöslau (Austria), Figs 54A 1 – A 3. NHMW 2010/0004/1648, SL: 2.1 mm, MD: 1.5 mm, Bad Vöslau (Austria), Figs 54B 1 –B 2. NHMW-ZOO-MO-75000-E-47001, SL: 3.5 mm, MD: 2.3 mm, Lăpugiu de Sus (Romania), Figs 54C 1 –C 3. Revised description. Shell small, relatively fragile, elongate ovate, with flattened spire (SL/MD = 1.6). Last whorl inflated, ovate, apex roundly truncated (MD/ApD = 2.7), whorl profile convex below, rounded at base. Periphery placed above mid-whorl (SL/ PH = 1.5). Sculpture of regular, chain-like spiral grooves; the links small, elongated and connected. Aperture total height of shell, moderately wide in upper quarter (MD/AW = 5.5), greatly widening anteriorly (MD/MAW = 1.2), resulting in high expansion rate (ER = 4.7); outer lip rising slightly above apex to slightly below, convex below, rounded abapically. Columella much reduced, narrow. Parietal callus very narrow. Discussion. See Landau et al. (2023: 399). Paleoenvironment. The species lives from the littoral zone in its more northern distribution to the bathyal zone in its southern distribution (Gofas et al. 2011). In the Paratethys it is known from shallow and deeper water environments. Distribution in Central Paratethys. Badenian (Middle Miocene): Vienna Basin: Bad Vöslau (Austria) (Berger 1953); Pannonian Basin: Szob (Hungary) (Strausz 1966); Făget Basin: Lăpugiu de Sus (Romania) (hoc opus). Proto-Mediterranean Sea. Serravallian (Middle Miocene): Karaman Basin: Akpınar, Lale, Seyithasan (Turkey) (Landau et al. 2013). Northeastern Atlantic. Tortonian (Late Miocene): Cacela Basin (Portugal) (Landau et al. 2023). North Sea. (Burdigalian/Langhian (Middle Miocene): the Netherlands (A.W. Janssen 1984). Pliocene to present-day. From North Sea to West Africa and Mediterranean Sea (see Landau et al. 2023: 399 for details).

Authors 3

1. Mathias Harzhauser

   Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655

2. Bernard Landau Aachen

   University of Lisbon · Instituto Dom Luiz · Naturalis Biodiversity Center

   Affiliation as printed

   Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494

3. Manuel António E. Malaquias

   University of Bergen · Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655 & Department of Natural History, University Museum of Bergen, University of Bergen, PB 7800, 5020 - Bergen, Norway manuel. malaquias @ uib. no; http: // orcid. org / 0000 - 0002 - 9668 - 945 X & Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655