Acteocina fusiformis

Zenodo (CERN European Organization for Nuclear Research)

Abstract

Acteocina fusiformis (Boettger, 1907) Figs 34A–J Bullina Lichtensteinii E. — Hauer 1837: 421 [non Acteocina lichtensteinii (Eichwald, 1830)]. * Tornatina fusiformis n. sp. —Boettger 1907: 203. Acteocina fusiformis (Boettger) — Zilch 1934: 277, pl. 22, fig. 19. Acteocina heraclitica n. sp. — Berger 1949: 83. Acteocina heraclitica n. sp. — Berger 1953: 106, pl. 18, figs 65–71. A [cteocina]. (A [cteocina].) heraclitica Berg.— Sieber 1958: 164. Acteocina heraclitica Berger— Švagrovský 1960: 109, pl. 14, figs 11–12. Acteocina (Acteocina) lajonkaireana Bast.— Florei 1961: 689, pl. 7, fig. 45 [non Acteocina lajonkaireana (de Basterot, 1825)]. Acteocina lajonkaireana Basterot— Strausz 1962: 153, pl. 74, figs 25–26 [non Acteocina lajonkaireana (de Basterot, 1825)]. Acteocina lajonkaireana Basterot, 1825 — Strausz 1966: 472, text-fig. 211a, pl. 74, figs 25–26 [non Acteocina lajonkaireana (de Basterot, 1825)]. Acteocina (Acteocina) lajonkaireana Bast.—Rado & Muţui 1970: 148, pl. 6, fig. 28 [non Acteocina lajonkaireana (de Basterot, 1825)]. Tornatina lajonkaireana (Bast.) — Urbaniak 1974: 40, pl. 4, fig. 9 [non Acteocina lajonkaireana (de Basterot, 1825)]. Acteocina lajonkaireana (Basterot, 1825) — Švagrovský 1984: 189, pl. 3, fig. 4 [non Acteocina lajonkaireana (de Basterot, 1825)]. Acteocina lajonkaireana (Basterot, 1825) —Harzhauser 2002: 127: pl. 11, fig. 18 [non Acteocina lajonkaireana (de Basterot, 1825)]. Acteocina lajonkaireana (Basterot, 1825) — Hladilová & Fordinál 2013: 39, fig. 5e [non Acteocina lajonkaireana (de Basterot, 1825)]. Acteocina lajonkaireana (de Basterot 1825) — Landau et al. 2013: 333, pl. 53, figs 3–4, pl. 77, fig. 9 [non Acteocina lajonkaireana (de Basterot, 1825)]. Acteocina heraclitica Berger, 1949 — Harzhauser et al. 2018a: 155, fig. 10T. non Acteocina fusiformis (Boettger, 1905 [sic])— Iljina 1993: 122, pl. 15, figs 15–16 [Acteocina sp.]. Type material. Lectotype (designated by Zilch 1934: 277): XII 2546 a, SL: 2.5 mm, MD: 1.1 mm, Coşteiu de Sus (Romania), illustrated in Zilch (1934: pl. 22, fig. 19). The specimen was destroyed during WW II and the photographic glass plate from Zilch (1934) is damaged. Illustrated material. NHMW 1876/0011/0170, SL: 2.7 mm, MD: 1.2 mm, Lăpugiu de Sus (Romania), Figs 34A 1 –A 2. NHMW 2020/0004/1661, SL: 3.8 mm, MD: 1.7 mm, Bad Vöslau (Austria), Figs 34B 1 –B 3. NHMW 2020/0004/1661a, SL: 3.3 mm, MD: 1.6 mm, Bad Vöslau (Austria), Figs 34C 1 –C 2. NHMW 1867/0019/0322, SL: 3.2 mm, MD: 1.5 mm, Coșteiu de Sus (Romania), Figs 34D 1 –D 2. NHMW 1997z0178/1210, SL: 3.8 mm, MD: 1.6 mm, Bad Vöslau (Austria), Figs 34E 1 –E 3. NHMW 1997z0178/1210a, SL: 3.2 mm, MD: 1.4 mm, Bad Vöslau (Austria), Figs 34F 1 –F 2. NHMW 1876/0011/0170a, SL: 4.1 mm, MD: 1.8 mm, Lăpugiu de Sus (Romania), Figs 34G 1 –G 3. NHMW 2008/0248/0479, SL: 2.4 mm, MD: 1.1 mm, Coșteiu de Sus (Romania), Figs 34H 1 –H 3. NHMW 1859/0037/0570, SL: 3.3 mm, MD: 1.5 mm, Vienna /Pötzleinsdorf (Austria), Figs 34I. NHMW-ZOO-MO-75000-E-3317A, SL: 3.4 mm, MD: 1.5 mm, Bad Vöslau (Austria), holotype of Acteocina heraclitica Berger, 1949, illustrated in Berger (1953: pl. 19, fig. 70), Fig. 34J. Revised description. Shell small, solid, moderately slender, elongate ovate with slightly convex sides (SL/MD = 2.2–2.3). Protoconch of about 1.5 whorls, flattened, strongly tilted (~140°), with nucleus two-thirds covered by first teleoconch whorl. Teleoconch of about three whorls separated by deeply canaliculated suture. Spire conical, weakly cyrtoconoid (apical angle 81–95°), broad (MD/ApD = 1.5–1.6). Early teleoconch whorls weakly convex, nearly straight sided with periphery at abapical suture, rapidly increasing in height. Last whorl 90–93% of total height, no shoulder, rounded at slowly contracting base. Periphery placed just above mid-whorl (SL/ PH = 1.7–1.8). Surface glossy with delicate, shallowly arcuate growth lines. Aperture 80–83% of total height, moderately narrow in upper half (MD/AW = 7.3–7.6), widening anteriorly (MD/MAW = 2.1–2.5), resulting in moderate expansion rate (ER = 3.0–3.5). Outer lip simple, thin, shallowly arcuate in profile, straight to slightly pinched inwards just below mid-whorl, rounded at base. Columella short, moderately to weakly excavated, thickened, forming narrow rim with prominent central columellar fold. Columellar callus delimiting narrow, shallow umbilical chink, persisting continuously into narrow, moderately thickened parietal callus. Paratethyan synonyms. Acteocina heraclitica Berger, 1949 is a subjective junior synonym of Acteocina fusiformis (Boettger, 1907). Berger (1949, 1953) did not discuss Acteocina fusiformis when describing Acteocina heraclitica and most probably overlooked this species, although he mentioned Acteocina heraclitica also from Coşteiu de Sus (Romania). Discussion. Acteocina fusiformis (Boettger, 1907) is a widespread species in the Badenian of the Central Paratethys Sea, which is characterized by its olivoid shell, without a shoulder, and by the prominent columellar fold. The northeastern Atlantic occurrences referred to as Acteocina lajonkaireana in the literature might be a species complex. Many specimens from the Early to Middle Miocene of France, described by Peyrot (1932), Glibert 1952a) and Lozouet et al. (2001) as Acteocina lajonkaireana agree with Acteocina fusiformis in their slender outline and columellar fold, whilst the syntype of Acteocina lajonkaireana is comparatively stout and has a weak fold (see Fig. 37H). Specimens from the Tarkhanian and Chokrakian of Russia, illustrated as this species by Iljina (1993), differ in their high spire, strongly convex spire whorls and incised but not canaliculate suture and represent another species. Paleoenvironment. Shallow marine, inner neritic; e.g., Bernhardsthal (Austria) in Harzhauser et al. (2018a). Distribution in Central Paratethys. Karpatian (Early Miocene): Korneuburg Basin: Kleinebersdorf, Stetten, Weinsteig (Austria) (Harzhauser 2002). Badenian (Middle Miocene): Polish Fore-Carpathian Basin: Błonie (Poland) (Urbaniak 1974). North Alpine Foreland Basin: Guntersdorf (Austria) (Berger 1953); Vienna Basin: Bernhardsthal, Bad Vöslau, Enzesfeld, Steinebrunn, Vienna /Pötzleinsdorf (Austria) (Berger 1953; Harzhauser et al. 2018a), Borský Mikuláš (Slovakia) (Švagrovský 1984); Danube Basin: Modra-Kráľová (Slovakia) (Hladilová & Fordinál 2013); Oberpullendorf Basin: Ritzing (Austria) (Berger 1953); Pannonian Basin: Várpalota (Hungary) (Strausz 1966); Eastern Slovakian Basin: Kuzmice (Slovakia) (Švagrovský 1960); Dacian Basin: Islaz (Romania) (Rado & Muţui 1970); Banat: Zorlenţul-Mare (Romania) (Florei 1961); Făget Basin: Coşteiu de Sus, Lăpugiu de Sus (Romania) (Boettger 1907). Proto-Mediterranean Sea. Serravallian (Middle Miocene): Karaman Basin: Seyithasan (Turkey) (Landau et al. 2013).

Authors 3

1. Mathias Harzhauser

   Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655

2. Bernard Landau Aachen

   University of Lisbon · Instituto Dom Luiz · Naturalis Biodiversity Center

   Affiliation as printed

   Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494

3. Manuel António E. Malaquias

   University of Bergen · Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655 & Department of Natural History, University Museum of Bergen, University of Bergen, PB 7800, 5020 - Bergen, Norway manuel. malaquias @ uib. no; http: // orcid. org / 0000 - 0002 - 9668 - 945 X & Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655