Ringiculocosta costata

Abstract

Ringiculocosta costata (Eichwald, 1830) Figs 12A–B, D, 13B * M [arginella]. costata m.— Eichwald 1830: 221. Marginella cancellata Nov.— Dubois de Montpéreux 1831: 24, pl. 1, figs 17–18. 1851 Ring [icula]. costata — Eichwald 1851: 125, pl. 10, fig. 44. Ringicula costata — Eichwald 1852: pl. 10, fig. 44. Ring [icula]. costata m.— Eichwald 1853: 259. Ringicula costata Eichw.—Hörnes 1852: 88, pl 9, figs 5a–d. Ringicula costata Eichw.— Hoernes & Auinger 1880: 70. [Ringicula costata Eichwald, 1830] var sicula n.— Seguenza 1881: 43, pl. 2, figs 9, 9a. [Ringicula costata Eichwald, 1830] var tenuis n.— Seguenza 1881: 43, pl. 2, fig. 10. Ringicula (Ringiculocosta) costata (Eichwald) — Cossmann 1895: 117, pl. 7, fig. 11. Ringicula costata Eichw. — Boettger 1902: 12. Ringicula costata Eichwald— Friedberg 1928: 559, pl. 36, figs 14–16. Ringicula (Ringiculocosta) costata (Eichwald) — Meznerics 1933: 351. Ringicula (Ringiculocosta) costata costata (Eichwald) — Berger 1954: 133, pl. 12, figs 64–67. Ringicula (Ringiculocosta) costata Eichw.— Strausz 1954: 38, pl. 5, fig. 118. Ringicula (Ringiculocosta) costata (Eichwald) — Korobkov 1955: pl. 113, figs 13–14. R [ingicula]. (R [ingiculocosta.) costata (Eichw.) — Sieber 1958: 163. Ringicula (Ringiculocosta) costata (Eichwald) — Zilch 1959: 20, fig. 57. Ringicula (Ringiculocosta) costata Eichwald, (1830) 1853— Strausz 1966: 471, pl. 36, figs 10–11 [not pl. 36, figs 12–13 = Ringiculocosta cancellarioides (Seguenza, 1881)]. Ringicula costata (Eichwald, 1830) — Zelinskaya et al. 1968: 233, pl. 52, figs 19–20. Ringicula costata Eichwald— Atanacković 1969: 217, pl. 14, fig. 6. Ringicula (Ringiculocosta) costata (Eichwald) — Bałuk 1970: 119, pl. 14, figs 10–11. Ringicula (Ringiculella) costata Eichw.—Rado & Muţui 1970: 148, pl. 6, fig. 30. Ringicula costata Eichwald— Jakubowski & Musial 1977: 115, pl. 16, figs 17–18. Ringicula (Ringiculocosta) costata (Eichwald, 1830) — Švagrovský 1984: 188, pl. 3, fig. 3. Ringicula (Ringiculocosta) costata (Eichwald, 1830) — Atanacković 1985: 186, pl. 42, figs 1–2. Ringicula costata Eichwald— Popa et al. 1997: 129, figs 10–11. Ringicula costata cancellarioides Seguenza— Kókay 1996: 458, pl. 4, fig. 7 [non Ringiculocosta cancellarioides (Seguenza, 1881)]. Ringicula (Ringiculocosta) costata (Eichwald) — Popa & Chira 1999: 402, pl. 4, fig. 6. Ringicula (Ringiculocosta) costata (Eichwald, 1830) — Stein et al. 2016: 166, pl. 73, figs 3–4 [? not figs 1–2 = Ringiculocosta sp.]. Ringicula costata costata (Eichwald, 1830) — Guzhov 2022a: 1126, pl. 6, figs 1–7. non Ringicula costata Eichwald 1830 — Krach 1981: 79, pl. 23, fig. 12 [= Ringiculocosta cancellarioides (Seguenza, 1881)]. non Ringicula (Ringiculocosta) costata (Eichwald, 1831 [sic])— Moths et al. 2010: 90, text-fig. 50 = [Ringiculocosta sp.]. Type material. Lectotype (designated by Guzhov 2022a: 1126), SL: 2.4 mm, MD: 1.6 mm, PSM, no. 3/469a, Zhukivtsi (Ukraine), illustrated in Guzhov 2022a: pl. 6, fig. 1). Illustrated material. NHMW 1855/0045/0036, SL: 2.2 mm, MD: 1.5 mm, Steinebrunn (Austria), Figs 12A 1 –A 2. NHMW 1863/0015/0616, SL: 2.5 mm, MD: 1.7 mm, Niederleis (Austria), Figs 12B 1 –B 2, 13B. NHMW 1863/0015/0616, SL: 2.0 mm, MD: 1.4 mm, Buituri (Romania), Figs 12D 1 –D 2. Revised description. Shell small, with globose last whorl, very broad (SL/MD: 1.4–1.5) with moderately high conical spire (AA= 58–64°). Protoconch paucispiral of about 1.5 whorls (dp = 400 μm). Junction with teleoconch marked by beginning of cancellate sculpture. Teleoconch of three whorls separated by narrowly impressed, linear suture. Spire whorls strongly convex with rounded shoulder. Sculpture of close-set axial ribs separated by interspaces of about same width, overrun by weaker spiral cords, resulting in cancellate pattern. Last whorl 72–75% of total height, with periphery just above mid-whorl (SL/ PH = 1.9), weakly convex to faintly concave subsutural ramp, rounded shoulder, inflated below; sculpture occasionally more prominent and wider spaced along shoulder; base broad conical, constricted. Aperture 58–61% of total height, moderately narrow, widening abapically (MD/MAW = 4.0–4.5). Outer lip thickened by convex labial varix, bearing low central palatal swelling; anal sinus broad, shallow, U-shaped; siphonal canal very short, open, broad, deeply notched at tip. Columella with two strongly elevated, robust folds abapically, parietal denticle prominent, trigonal. Columellar and parietal callus strongly thickened, sharply delimited, narrowly expanded over base. Discussion. Ringiculocosta costata is quite variable concerning its sculpture, which led Seguenza (1881) to introduce the varieties tenuis (finely cancellate) and sicula (coarsely cancellate). Paratethyan specimens display a comparable variability within the same locality and taxonomic separation of the various morphs is unjustified. The situation seems to be different in the Eastern Paratethys Sea from where Guzhov (2022a) described three subspecies of Ringiculocosta costata from the Tarkhanian and Chokrakian (Middle Miocene). These taxa seem to represent a distinct radiation in the largely isolated Eastern Paratethys Sea. Ringiculocosta plicatula (Mayer, 1871), from the Burdigalian (Early Miocene) of Saucats (France), differs in its slenderer shell, higher last whorl, weaker shoulder and predominant axial sculpture (see Mayer 1871: 347, pl. 9, fig. 8; Morlet 1878: 267, pl. 6, fig. 9). Similarly, Ringiculocosta cancellarioides (Seguenza, 1881) differs in its slenderer outline. Paleoenvironment. Shallow marine, inner neritic based on associated fauna (own data, M.H.). Distribution in Central Paratethys. Badenian (Middle Miocene): Polish-Ukrainian Fore-Carpathian Basin: Borki Wielkie (Poland), Hołdy, Hryts’kiv, Faszczówka, Tarnoruda, Ripyntsi, Shushkivtsi, Staryi Pochaiv, Zboriv, Zalistsi, Zhabyak, Zhukivtsi, Varivtsi, Yaseniv (Ukraine) (Friedberg 1938; Guzhov 2022a); Voronyaky Hills: Golubytsya (Hołubica), Pidgirtsi (Ukraine); North Alpine-Carpathian Foreland Basin: Lysice (Czech Republic) (Berger 1954); Vienna Basin: Bad Vöslau, Niederleis, Steinebrunn, Vienna /Grinzing (Austria) (Berger 1954), Mikulov (Czech Republic) (Hörnes 1852), Borský Mikulás (Slovakia) (Švagrovský 1984); Eisenstadt-Sopron Basin: Forchtenau (Austria) (Berger 1954); Oberpullendorf Basin: Ritzing (Austria) (Berger 1954); Pannonian Basin: Budapest, Hidas (Hungary) (Strausz 1966; Kókay 1996); Southern Pannonian Basin: Hrvaćani (Bosnia and Herzegovina) (Atanacković 1985); Făget Basin: Coşteiu de Sus (Romania) (Boettger 1902); Dacian Basin: Islaz (Romania) (Rado & Muţui 1970); Borod Basin: Cetea Valley (Romania) (Popa & Chira 1999). Proto-Mediterranean Sea. Tortonian (Late Miocene): Po Basin: Montegibbio (Italy) (Sacco 1892); Emilia-Romagna: Modena (Italy), Sicily: Ciminna, Rometta (Italy) (Seguenza 1881). Atlantic. Tortonian (Late Miocene): Loire Basin, NW France (BL unpublished data). North Sea. Hemmoorian (Early Miocene): North Sea Basin: Germany (Stein et al. 2016).

Authors 3

1. Mathias Harzhauser

   Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655

2. Bernard Landau Aachen

   University of Lisbon · Instituto Dom Luiz · Naturalis Biodiversity Center

   Affiliation as printed

   Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494

3. Manuel António E. Malaquias

   University of Bergen · Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655 & Department of Natural History, University Museum of Bergen, University of Bergen, PB 7800, 5020 - Bergen, Norway manuel. malaquias @ uib. no; http: // orcid. org / 0000 - 0002 - 9668 - 945 X & Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655