Ringicula minor

Abstract

Ringicula minor (Grateloup, 1838) Figs 14A–E * Auricula ringens var. b. minor — Grateloup 1838: 260, 286, pl. 6, fig. 8. Ringicula buccinea Desh.—Grateloup 1846: pl. 11, fig. 8–10 [non Ringicula buccinea (Brocchi, 1814)]. Ringicula buccinea Desh.—Hörnes 1852: 86 (pars), pl. 9, fig. 4 [non Ringicula buccinea (Brocchi, 1814)]. Ringicula Paulucciae Morlet— Morlet 1878: 266, pl. 6, fig. 6, pl. 8, fig. 9. Ringicula (Ringiculella) Tournoueri L. Morlet— Morlet 1878: 287, pl. 6, fig. 10. Ringicula auriculata Men.— Friedberg 1928: 551, pl. 36, figs 9–11 [non Ringicula auriculata (Ménard de la Groye, 1811)]. Ringicula (Ringiculella) Tournoueri Morlet— Peyrot 1932: 143, no 1371 ‘bis’, pl. 11, figs 7, 11–13, 18–25, 37, 54, 61. Ringicula (Ringiculella) auriculata paulucciae Morlet— Berger 1954: 115, pl. 7, figs 3–18. Ringicula (Ringiculella) auriculata elongata Morlet— Berger 1954: 120, pl. 8, figs 33–36 [non Ringicula elongata Morlet, 1878]. Ringicula (Ringiculella) auriculata buccinea Brocchi, 1814 — Strausz 1954: 38, pl. 5, fig. 117 [non Ringicula buccinea (Brocchi, 1814)]. Ringicula auriculata Menardi— Moisescu 1955: 175, pl. 15, figs 8–9 [non Ringicula auriculata (Ménard de la Groye, 1811)]. R [ingicula]. (R [ingiculina].) auriculata elongata Morl.— Sieber 1958: 162 [non Ringicula elongata Morlet, 1878]. R [ingicula]. (R [ingiculina].) paulucciae Morl.— Sieber 1958: 162. ? Ringicula auriculata buccinea Brocchi— Boda 1959: 724, pl. 32, fig. 4 [non Ringicula buccinea (Brocchi, 1814)]. Ringicula (Ringiculella) auriculata buccinea Br. — Csepreghy-Meznerics 1956: 436, pl. 12, figs 13–14 [non Ringicula buccinea (Brocchi, 1814)]. Ringicula (Ringiculina) auriculata Ménard var.— Strausz 1962: 91, pl. 36, figs 8–9 [non Ringicula auriculata (Ménard de la Groye, 1811)]. Ringicula auriculata paulucciae Morlet 1878 — Steininger 1963: 56, pl. 12, fig. 15. Ringicula (Ringiculina) auriculata Ménard var.— Strausz 1966: 471, pl. 36, figs 8–9 [non Ringicula auriculata (Ménard de la Groye, 1811)]. Ringicula auriculata paulucciae Morlet— Kókay 1966: 66, pl. 10, fig. 11. Ringicula (Ringiculella) paulucciae Morlet, 1878 — Švagrovský 1984: 184, pl. 2, fig. 6. Ringicula (Ringiculella) elongata Morlet, 1878 — Švagrovský 1984: 185, pl. 3, fig. 1 [non Ringicula elongata Morlet, 1878]. Ringicula (Ringiculella) laevigata (Eichwald, 1830) — Švagrovský 1984: 186, pl. 3, fig. 2 [non Ringicula laevigata (Eichwald, 1830)]. Ringicula (Ringiculella) auriculata buccinea (Brocchi, 1814) — Atanacković 1985: 186 (pars), pl. 41, figs 13–14 [non Ringicula buccinea (Brocchi, 1814)]. Ringicula paulucciae Morlet— Valdés & Héros 1998: 700, fig. 2f. Ringicula tournoueri Morlet— Valdés & Héros 1998: 700, fig. 3g. Ringicula minor (Grateloup) — Lozouet et al. 2001: 80, pl. 37, fig. 1. Ringicula (Ringiculella) auriculata paulucciae Morlet, 1878 —Harzhauser 2002: 125, pl. 11, fig. 16. Ringicula minor (Grateloup, 1828 [sic])— Lesport & Cahuzac 2005: 98, pl. 4, figs 7–8. Ringicula minor (Grateloup, 1838) — Harzhauser et al. 2011: 218, fig. 6.2. Ringicula minor (Grateloup, 1838) — Cahuzac et al 2012: 403, pl. 2B, fig. 6. Ringicula minor (Grateloup, 1838) — D’Amico et al. 2012: 170, pl. 3, fig. 36. Ringicula minor (Grateloup, 1838) — Landau et al. 2013: 326, pl. 52, fig. 16, pl. 76, fig. 14, pl. 77, fig. 1. Ringicula minor (Grateloup, 1838) — Thivaiou et al. 2019: 346, fig. 7C1, C2. Ringicula minor (Grateloup, 1838) — Dominici et al. 2020: 85, fig. 12I. Ringicula minor (Grateloup, 1838) — Landau et al. 2020: pl. 13, figs 1–2. Type material. Syntype illustrated in Grateloup (1838: 286, pl. 6, fig. 8), Aquitaine Basin, no type locality given, Early Miocene. Illustrated material. NHMW-ZOO-MO-75000-E-57962, SL: 5.8 mm, MD: 3.7 mm, Bad Vöslau (Austria), Figs 15E 1 –E 2. NHMW 1862/0001/0521, SL: 6.1 mm, MD: 3.8 mm, Buituri (Romania), Figs 14A 1 –A 2. NHMW 1862/0001/0521a, SL: 6.0 mm, MD: 3.6 mm, Buituri (Romania), Figs 14B 1 –B 2. NHMW 1850/0002/0012, SL: 5.0 mm, MD: 3.1 mm, Grund (Austria), Figs 14C 1 –C 2. NHMW 1862/0001/0521b, SL: 4.9 mm, MD: 2.9 mm, Buituri (Romania), Figs 14D 1 –D 2. Revised description. Shell medium sized, moderately globose, broad (SL/MD = 1.5–1.6), with high conical spire (AA= 58–62°). Protoconch unknown. Teleoconch of four whorls separated by narrowly impressed, linear suture. Spire whorls moderately convex with periphery placed some distance above abapical suture. Sculpture of distinct, broad spiral cords separated by narrow spiral grooves. Last whorl 70–73% of total height, globose, with periphery placed mid-whorl (SL/ PH = 2.0–2.1); weakly constricted at base, glossy, with numerous, broad, flattish spiral cords separated by narrow grooves. Aperture 64–66% of total height, adapically moderately narrow, widening abapically (MD/MAW = 3.6–4.6). Outer lip very strongly thickened by wide labial varix bearing broad palatal swelling. Anal sinus broad, moderately deep, U-shaped; siphonal canal very short, open, broad, notched at tip. Columella with two prominent, robust folds abapically, parietal denticle prominent or weak; fused with broad parietal pad. Columellar and parietal callus thickened, sharply delimited, slightly expanded over base. Discussion. Ringicula minor (Grateloup, 1838) differs from Ringicula guzhovi Landau, Harzhauser & Malaquias, 2023 in its higher spire, less inflated last whorl and prominent spiral sculpture. See Landau et al. (2013: 326) for further discussion. Paleoenvironment. Shallow marine, inner neritic (own data M.H). Distribution in Central Paratethys. Eggenburgian (Early Miocene): North Alpine Foreland Basin: Fels am Wagram (Austria) (Steininger 1963). Karpatian (Early Miocene): North Alpine-Carpathian Foreland Basin: Laa (Austria) (Berger 1954), Malé Karpaty Mts.: Cerová (Slovakia) (Harzhauser et al. 2011); Korneuburg Basin: Stetten (Austria) (Harzhauser 2002). Badenian (Middle Miocene): North Alpine-Carpathian Foreland Basin: Kalladorf, Grund, Guntersdorf (Austria) (Berger 1954); Badenian (Middle Miocene): Voronyaky Hills: Oles’ko (Ukraine) (Friedberg 1928); Korytnica Basin: Korytnica (Poland) (Friedberg 1928); Vienna Basin: Borský Mikuláš (Czech Republic) (Švagrovský 1984); Bad Vöslau, Steinebrunn, Vienna /Grinzing, Vienna /Pötzleinsdorf (Austria) (Berger 1954); Boršov (Porstendorf), Mikulov, Jaroměřice (Czech Republic) (Berger, 1954); Oberpullendorf Basin: Ritzing (Austria) (Berger 1954); Pannonian Basin: Herend-Márkó, Letkés, Várpalota, Szob (Hungary) (Strausz 1954; Csepreghy-Meznerics 1956; Kókay 1966); Southern Pannonian Basin: Miljevići (Bosnia and Herzegovina) (Atanacković 1985); Făget Basin: Buituri (Romania) (hoc opus). Proto-Mediterranean Sea. Serravallian (Middle Miocene): Karaman Basin: Akpınar, Lale, Seyithasan (Turkey) (Landau et al. 2013). Tortonian (Late Miocene): Livorno Hills: Quarata (Italy) (Dominici et al. 2020); Calabria: Cessaniti (Italy) (D’Amico et al. 2012). Northeastern Atlantic. Burdigalian (Early Miocene): Aquitaine Basin: Cestas, Léognan, Martillac, Mérignac, Saucats, Pourquey, Peloua, Saint-Paul-lès-Dax (France) (Peyrot 1932; Lesport & Cahuzac 2005). Serravallian (Middle Miocene): Salles, Sallespisse, Salies-de-Béarn (France) (Peyrot 1932). Tortonian (Late Miocene): Loire Basin (France) (Landau et al. 2020).

Authors 3

1. Mathias Harzhauser

   Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655

2. Bernard Landau Aachen

   University of Lisbon · Instituto Dom Luiz · Naturalis Biodiversity Center

   Affiliation as printed

   Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494

3. Manuel António E. Malaquias

   University of Bergen · Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655 & Department of Natural History, University Museum of Bergen, University of Bergen, PB 7800, 5020 - Bergen, Norway manuel. malaquias @ uib. no; http: // orcid. org / 0000 - 0002 - 9668 - 945 X & Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655