Pyrunculus elongatus

Abstract

Pyrunculus elongatus (Eichwald, 1830) Figs 19A–E * Bulla elongata m.— Eichwald 1830: 214. Bulla lignarioides — Andrzejowski 1833: 446, pl. 11, fig. 4. Bulla elongata m.— Eichwald 1851: 145, pl. 11, figs 15a–c. Bulla elongata — Eichwald 1852: pl. 11, figs 15a–c. Bulla elongata m.— Eichwald 1853: 304. Bulla conulus Desh.—Hörnes 1856: 620, pl. 50, figs 4a–c [non Pyrunculus conulus (Deshayes, 1824)]. B [ulla]. conulus Desh.— Reuss 1867: 161 [non Pyrunculus conulus (Deshayes, 1824)]. Cylichnina elongata (Eichw.) — Boettger 1902: 178. Cylichnina elongata (Eichw.) —Boettger 1907: 206. Bullinella elongata Eichw.— Friedberg 1928: 553, pl. 36, fig. 6. Bullinella (Cylichnina) elongata (Eichwald) — Meznerics 1933: 351. Retusa (Cylichnina) elongata (Eichwald) — Berger 1953: 97 (pars), pl. 17, figs 33–35. Retusa (Cylichnina) elongata Eichw. — Csepreghy-Meznerics 1956: 423, pl. 12, figs 15–18. R [etusa]. (C [ylichnina]). elongata (Eichw.) — Sieber 1958: 163. Retusa (Cylichnina) elongata (Eichw.) — Báldi 1960: 90. Retusa (Cylichnina) elongata Eichwald— Strausz 1966: 474, text-fig. 212a, pl. 74, figs 30, 31. Cylichnina elongata (Eichwald, 1830) — Zelinskaya et al. 1968: 235, pl. 52, figs 29–30. Pyrunculus elongatus (Eichwald, 1830) — Landau et al. 2013: 338 (pars), pl. 53, fig. 11 [non pl. 53, fig. 10, pl. 77, fig. 12 = Pyrunculus parangistoma (Boettger, 1902)]. non Bullinella (Cylichnina) elongata Eichwald— Peyrot 1932: 190, no. 1399, pl. 13, figs 53, 54 [= see discussion]. non Retusa (Cylichnina) elongata Eichwald— Glibert 1952b: 143, pl. 10, fig. 18 [= see discussion]. non Retusa (Cylichnina) elongata (Eichwald 1830) —Rasmussen 1956: 103, pl. 10, fig. 3 [= see discussion]. non Retusa elongata (Eichwald) — Sorgenfrei 1958: 345, pl. 74, fig. 254 [= see discussion]. non Retusa elongata (Eichwald 1830) — Anderson 1964: 335, pl. 51, fig. 303 [= see discussion]. non Retusa (Pyrunculus) elongata (Eichwald, 1830) — Nordsieck 1972: 124, pl. 32, fig. 217 [= see discussion]. non Retusa (Cylichnina) elongata (Eichwald, 1830) —A.W. Janssen 1984: 378, pl. 19, figs 10–11 [= see discussion]. non Pyrunculus elongatus (Eichwald, 1850) — Švagrovský 1984: 192, pl. 3, fig. 7 [= Pyrunculus leuchsi (Berger, 1949)]. non Cylichnia [sic] elongata (v. Eichwald, 1830)— Moths 1989: 112, pl. 22, fig. 120 [= see discussion]. non Retusa (Cylichnina) elongata (Eichwald, 1830) — Iljina 1993: 125, pl. 15, figs 29–31 [= see discussion]. non Cylichnina elongata (Eichwald, 1830) — Harzhauser & Kowalke 2002: 74, pl. 13, fig. 17 [= Retusidae indet.]. non Retusa (Cylichnina) elongata (Eichwald, 1830) — Wienrich 2007: 778, pl. 134, figs 1–2, pl. 172, figs 3–4 [= see discussion]. non Cylichnina elongata (Eichwald, 1830) — Moths et al. 2010: 92, pl. 25, fig. 11 [= see discussion]. non Pyrunculus elongatus (Eichwald, 1830) — Landau et al. 2013: 338 (pars), pl. 53, fig. 10, pl. 77, fig. 12 [= Pyrunculus parangistoma (Boettger, 1902)]. Type material. Lectotype (designated herein): specimen illustrated in Eichwald (1851: pl. 11, figs 15a–c). probably from Żukowce (Zhukivtsi) (or Tarnoruda, Zaleśce) (Ukraine). Probably stored in the St. Petersburg State University, Paleontological Museum (Russia). Illustrated material. NHMW 2010/0004/1656, SL: 5.7 mm, MD: 2.3 mm, Bad Vöslau (Austria), Figs 19A 1 –A 3. NHMW 2010/0004/1656a, SL: 6.0 mm, MD: 2.2 mm, Bad Vöslau (Austria), Figs 19B 1 –B 3. NHMW 1869/0001/0775, SL: 4.8 mm, MD: 2.0 mm, Coşteiu de Sus (Romania), Figs 19C 1 –C 2. NHMW 2010/0004/1656b, SL: 5.1 mm, MD: 2.2 mm, Bad Vöslau (Austria), Figs 19D 1 –D 2. NHMW-ZOO-MO-75000-E-58763, SL: 6.6 mm, MD: 2.5 mm, Gainfarn (Austria), Figs 19E 1 –E 3. Revised description. Shell small, medium-thickness, slender (SL/MD = 2.5–2.6), elongate pyriform, only last whorl visible. Last whorl with almost straight sides, forming angle of about 19–21°. Periphery placed close to one-third whorl height (SL/ PH = 3.0–3.6), rounded at moderately contracting base. Apex moderately narrow (MD/ApD = 2.5–2.6), truncated, apical plane slightly oblique (AP = 12–15°), deeply and narrowly umbilicate, delimited by round-edged, faintly swollen, elevated crest. Sculpture of a few, faint spiral grooves over base. Aperture longer than shell, very narrow down to periphery (MD/AW = 13.8–15.7), abruptly widening anteriorly below periphery (MD/ MAW = 1.9–2.2), resulting in high expansion rate (ER = 6.6–8.0). Outer lip thin, slightly alate apically, extending some distance above apex, weakly concave just above periphery. Columella short, with narrow but prominent columellar callus with low, broad columellar fold. Columellar callus short, narrow, delimiting deep umbilical chink. Parietal callus not developed. Discussion. This is a very distinctive species due to its slender conical outline and weak columellar fold. Nevertheless, the name was erroneously used for many Miocene shells, which in our opinion are not conspecific with Pyrunculus elongatus (Eichwald, 1830). Middle Miocene specimens from the northeastern Atlantic and North Sea illustrated by Peyrot (1932), Glibert (1952b), A.W. Janssen (1984) and many others listed above are comparatively stout with more convex whorls. We have not reviewed this group but note that Pyrunculus subconulus (d’Orbigny, 1852) might be an available name for at least some of these specimens. A comment on Pyrunculus hoernesi (Weinkauff, 1866). Weinkauff (1866: 238) established Bulla (Cylichna) hoernesi when describing extant specimens from Algeria but referred to the illustration of a Middle Miocene specimen from the Vienna Basin illustrated in Hörnes (1856: pl. 50, figs 4a–b) as Bulla conulus Deshayes, 1824 [= Pyrunculus elongatus (Eichwald, 1830)]. Thus, as pointed out by Tringali & Oliverio (2001: 157), this Miocene specimen belongs to the type material of Bulla (Cylichna) Hoernesi following ICZN 1999 Art., 72.4. No lectotype was designated by Tringali & Oliverio (2001) from the Algerian shells and therefore the status of Pyrunculus hoernesi (Weinkauff, 1866) is still problematic. Choosing the Paratethyan specimen as lectotype would make Bulla hoernesi Weinkauff, 1866 a subjective junior synonym of Bulla elongata Eichwald, 1830. Therefore, we select the syntype stored in the Zoological Museum Rome (Italy), Monterosato coll. box 16118, as lectotype. The specimen was illustrated in Tringali & Oliverio (2001: fig. 16), SL: 2.8 mm, MD: 1.3 mm, Alger (Algeria). Paleoenvironment. Occurs in inner neritic to outer neritic assemblages. Distribution in Central Paratethys. Badenian (Middle Miocene): Voronyaky Hills: Golubytsya (Hołubica), Yaseniv (Friedberg 1938); Korytnica Basin: Korytnica (Poland) (Friedberg 1938); Polish-Ukrainian Fore-Carpathian Basin: Wieliczka (Poland) (Reuss 1867); Fashchivka, Staryi Pochaiv, Tarnoruda, Zaleśce (Żabiak), Zhukivtsi (Ukraine) (Friedberg 1938); Vienna Basin: Baden, Gainfarn, Steinebrunn, Vienna /Pötzleinsdorf (Austria) (Bergen 1953); Pannonian Basin: Letkés, Szob, Szokolya (Hungary) (Báldi 1960; Strausz 1966); Făget Basin: Coşteiu de Sus, Lăpugiu de Sus (Romania) (Boettger 1907). Proto-Mediterranean Sea. Serravallian (Middle Miocene): Karaman Basin: Seyithasan (Turkey) (Landau et al. 2023).

Authors 3

1. Mathias Harzhauser

   Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655

2. Bernard Landau Aachen

   University of Lisbon · Instituto Dom Luiz · Naturalis Biodiversity Center

   Affiliation as printed

   Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494

3. Manuel António E. Malaquias

   University of Bergen · Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655 & Department of Natural History, University Museum of Bergen, University of Bergen, PB 7800, 5020 - Bergen, Norway manuel. malaquias @ uib. no; http: // orcid. org / 0000 - 0002 - 9668 - 945 X & Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655