Retusa truncatula

Zenodo (CERN European Organization for Nuclear Research)

Abstract

Retusa truncatula (Bruguière, 1792) Figs 30A–E * Bulla truncatula Nob.—Bruguière 1792: 377. Retusa truncatula (Brug.) —Boettger 1907: 204. Tornatina truncatula Brug.— Friedberg 1928: 544, pl. 36, figs 1–2. Tornatina truncatula Brug.— Friedberg 1938: 160. Bulla truncatula Brug.— Jekelius 1944: 91, pl. 26, figs 13–16. Retusa (Retusa) truncatula grundensis n. subsp. — Berger 1949: 80 [nomen nudum]. Retusa (Retusa) truncatula subtruncatula (Orbigny) — Berger 1953: 89, pl. 16, fig. 15. Retusa (Retusa) truncatula grundensis n. subsp. — Berger 1953: 90, pl. 16, fig. 16. Retusa (Retusa) truncatula truncatula (Bruguière) — Berger 1953: 88, pl. 16, figs 8–14, text-figs 2, 3, 5. Retusa (Retusa) truncatula pappi n. subsp. — Berger 1953: 91, pl. 16, fig. 19. Retusa (Retusa) truncatula pappi Berger— Papp 1954: 58, pl. 10, fig. 1. R [etusa]. (R [etusa].) t. pappi Berg.— Sieber 1958: 163. R [etusa]. (R [etusa].) t. subtruncatula (Orb.) — Sieber 1958: 163. R [etusa]. (R [etusa].) t. grundensis Berg.— Sieber 1958: 163. R [etusa]. (R [etusa].) truncatula truncata (Ad.) — Sieber 1958: 163. ? Retusa truncatula (Adams) — Boda 1959: 725, pl. 32, fig. 5. Retusa truncatula pappi Berger— Švagrovský 1964: 84, pl. 21, fig. 5. Retusa truncatula Bruguière, 1792 — Strausz 1966: 474, pl. 74, figs 27–28. Retusa truncatula (Bruguière, 1792) — Zelinskaya et al. 1968: 234, pl. 52, figs 23–24. Retusa truncatula (Bruguière) — Bałuk 1970: 119, pl. 14, fig. 7. Retusa truncatula pappi Berger, 1953 — Švagrovský 1971: 419, pl. 72, figs 7–9. Retusa (Retusa) truncatula Brug.—Rado & Muţui: 148, pl. 6, figs 14–16, 23. Retusa truncatula (Bruguière, 1792) — Ionesi & Ionesi 1972: pl. 4, figs 6–10. Retusa truncatula truncatula (Bruguière 1792) — Krach 1981: 79, pl. 23, fig. 17. Retusa (Coleophysis) truncatula (Bruguière, 1792) — Švagrovský 1984: 191, pl. 3, fig. 6. Retusa (Retusa) truncatula (Bruguière) — Atanacković 1985: 190, pl. 42, figs 13–14. Retusa truncatula (Bruguière, 1792) — Lukeneder et al. 2011: 772, fig. 4X. Retusa truncatula (Bruguière, 1792) — Tămaș et al. 2013: 78, fig. 4h. Retusa truncatula (Bruguière 1792) — Chirli 2013: 49, pl. 11, figs 11–18. Retusa truncatula (Bruguière, 1792) — Landau et al. 2013: 337, pl. 77, fig. 11. Retusa truncatula (Bruguière, 1792) — Ceulemans et al. 2018: 118, pl. 7, fig. 8. Retusa sp. — Harzhauser et al. 2018b: 30, fig. 3/13. Retusa truncatula (Bruguière, 1792) — Landau et al. 2020: 268, pl. 14, fig. 1. Retusa truncatula (Bruguière, 1792) — Dominici et al. 2020: 85, fig. 12K. Retusa truncatula (Bruguière, 1792) — Guzhov 2022b: 1239, pl. 36, figs 9–10, pl. 37, figs 1–8 (cum syn.). Retusa truncatula (Bruguière, 1792) — Landau et al. 2023: 268, pl. 34, figs 1–2 (cum syn). non Retusa (Retusa) truncatula (Brugière) [sic]—Kojumdgieva in Kojumdgieva & Strachimirov 1960, p. 221, pl. 52, figs 7–8 [fig. 7 = Pyrunculus parangistoma (Boettger, 1902), fig. 8 = Pyrunculus sp.]. non Retusa (Retusa) truncatula (Bruguière) — Atanacković 1969: 217, pl. 14, fig. 4 [= Retusa sp.]. non Retusa truncatula (Bruguière, 1792) —Harzhauser 2002: 126, pl. 11, fig. 19 [= Retusa sp.]. Type material. We are not aware of the whereabouts of the type material. Illustrated material. NHMW 2018/0248/0476, SL: 2.9 mm, MD: 1.4 mm, Coșteiu de Sus (Romania), Figs 30A 1 –A 3. NHMW-ZOO-MO-75000-E-41686, SL: 3.9 mm, MD: 1.8 mm, Steinebrunn (Austria), illustrated as Retusa subtruncatula in Berger (1953: pl. 16, fig. 15), Figs 30B 1 –B 2. NHMW-ZOO-MO-75000-E-47002, SL: 3.1 mm, MD: 1.5 mm, Grund (Austria), Badenian, holotype of Retusa grundensis Berger, 1953, Figs 30C 1 –C 3. NHMW 2025/0152/0001, SL: 1.8 mm, MD: 0.9 mm, Bad Vöslau (Austria), Figs 30D 1 –D 3. NHMW 2025/0152/0002, SL: 1.3 mm, MD: 1.1 mm, Bad Vöslau (Austria), Figs 30E 1 –E 3. Revised description. See Landau et al. (2023: 384). Paratethyan synonyms. Retusa truncatula pappi Berger, 1953. Holotype, NHMW-ZOO-MO-75000-E-47018, Hölles (Austria), illustrated in Berger (1953: pl. 16, fig. 19, the specimen is lost. Retusa truncatula pappi was introduced by Berger (1953) for specimens with slight concavity below the shoulder, which does not justify a separation from Retusa truncatula. Specimens treated by Papp (1954) as Retusa truncatula pappi support this synonymization. For a list of synonyms based on extant specimens see MolluscaBase eds. (2025a). Discussion. For a discussion on the variability of this species in the Paratethys Sea see Guzhov (2022b: 1239). Retusa truncatula (Bruguière, 1792) is extremely similar to Retusa mystica sp. nov. but differs in usually being more cylindrical and having a pinched region just above the periphery. Moreover, when seen in apical view, the axial folds or riblets are present at the rounded shoulder that are not, or hardly, present in R. mystica sp. nov. Paleoenvironment. This species lives in sandy and muddy habitats up to 50 m depth (Gofas et al. 2011). Distribution in Central Paratethys. Badenian (Middle Miocene): Roztocze Hills: Łychów (Poland) (Krach 1981); Korytnica Basin: Korytnica (Poland) (Friedberg 1938); Nowy Sącz Basin: Niskowa (Poland) (Bałuk 1970); Voronyaky Hills: Oles’ko, Pidgirtsi, Yaseniv (Ukraine) (Friedberg 1938); Polish-Ukrainian Fore-Carpathian Basin: Wieliczka (Poland) (Reuss 1867); Borki Wielkie (Poland); Tarnoruda, Staryi Pochaiv, Zaleśce, Zhabyak (Ukraine) (Friedberg 1938); Korytnica Basin: Korytnica (Poland) (Friedberg 1938); Nowy Sącz Basin: Niskowa (Poland) (Bałuk 1970); North Alpine-Carpathian Foreland Basin: Grund, Guntersdorf (Austria) (Berger 1953): Vienna Basin: Baden, Bernhardsthal, Steinebrunn (Austria) (Berger 1953; Harzhauser et al. 2018a), Borský Mikuláš (Slovakia) (Švagrovský 1984); Pannonian Basin: Várpalota (Hungary) (Strausz 1966); Făget Basin: Coşteiu de Sus (Romania) (Boettger 1907); Eastern Slovakian Basin: Kuzmice (Slovakia) (Švagrovský 1960); Dacian Basin: Islaz (Romania) (Rado & Muţui 1970). Sarmatian (Middle Miocene): Polish-Ukrainian Fore-Carpathian Basin: Krzemieniec Żolobki (Poland); Rydomyl’, Shushkivtsi, Zalistsi (Ukraine) (Friedberg 1938); Vienna Basin: Hölles, Nexing (Austria) (Berger 1953; hoc opus), Vienna /Theresiengasse (Austria) (Harzhauser et al. 2018b); Holič, Podivín, Rohožník (Slovakia) (Švagrovský 1971); Eisenstadt-Sopron Basin: Wiesen (Austria) (Berger 1953); Pannonian Basin: Sopron, Pécsvárad, Karád (Hungary) (Boda 1959); Eastern Slovakian Basin: Slanec, Trstené pri Hornáde, Nižná Myšľa, Slančík, Bidovce, Sečovce (Slovakia) (Švagrovský 1971); Transylvanian Basin: Soceni (Romania) (Jekelius 1944); Poiana Ruscă Mountains: Răcăştia (Romania) (Tămaș et al. 2013). Eastern Paratethys. Widespread in the Volhynian and Bessarabian of Romania, Moldova and Russia (Guzhov 2022b). Proto-Mediterranean Sea. Serravallian (Middle Miocene): Karaman Basin:Akpınar, Lale, Seyithasan (Turkey) (Landau et al. 2013). Tortonian (Late Miocene): Livorno Hills: Quarata (Italy) (Dominici et al. 2020). Tortonian and Messinian (late Miocene): Po Basin (Italy) (Sacco 1897; Venzo & Pelosio 1963; d’Amico et al. 2012). Pliocene to present-day. North Sea, northeastern Atlantic and Mediterranean Sea (see Landau et al. 2023 for details).

Authors 3

1. Mathias Harzhauser

   Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655

2. Bernard Landau Aachen

   University of Lisbon · Instituto Dom Luiz · Naturalis Biodiversity Center

   Affiliation as printed

   Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494

3. Manuel António E. Malaquias

   University of Bergen · Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655 & Department of Natural History, University Museum of Bergen, University of Bergen, PB 7800, 5020 - Bergen, Norway manuel. malaquias @ uib. no; http: // orcid. org / 0000 - 0002 - 9668 - 945 X & Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655