Roxania callifera

Abstract

Roxania callifera (Boettger, 1907) Figs 52A–B Bulla utricula Brocc.—Hörnes 1856: 618 (pars), pl. 50, figs 2a–c [non Roxania utriculus (Brocchi, 1814)]. * Sabatia callifera n. sp. —Boettger 1907: 205. Sabatia callifera Boettger— Zilch 1934: 278, pl. 22, fig. 20. Roxania (Sabatia) callifera callifera (Boettger) — Berger 1953: 112, pl. 18, fig. 78. R [oxania]. (S [abatia].) callifera callifera (Bttg.) — Sieber 1958: 164. non Roxania (Roxania) callifera (Boettger, 1906) —Kojumdgieva in Kojumdgieva & Strachimirov 1960: 222, pl. 52, figs 10–11 [= Roxania utriculus (Brocchi, 1814)]. non Roxania (Roxania) califera [sic] (Boettger)— Atanacković 1969: 217, pl. 14, figs 5–5a [= Roxania utriculus (Brocchi, 1814)]. non Roxania (Roxania) callifera (Boettger, 1906) — Atanacković 1985: 188, pl. 42, figs 4–5 [= Roxania utriculus (Brocchi, 1814)]. non Sabatia callifera Boettger, 1906 — Harzhauser et al. 2011: 218, fig. 7.1 [= Roxania utriculus (Brocchi, 1814)]. Type material. Lectotype (designated by Zilch 1934: 278), SMF 363162 - 12.2548a (glass plate 401), SL: 4.8 mm, MD: 2.9 mm, Coşteiu de Sus (Romania), illustrated in Zilch (1934: pl. 22, fig. 20). The specimen was destroyed during WW II. Illustrated material. NHMW 1847/0037/0131, SL: 4.4 mm, MD: 2.9 mm, Baden (Austria), illustrated in Hörnes (1856: pl. 50, fig 2a–c), Figs 52A 1 –A 2. NHMW-ZOO-MO-75000-E-47090, SL: 4.2 mm, MD: 2.6 mm, Bad Vöslau (Austria), Figs 52B 1 –B 2. Revised description. Shell small to medium sized, solid, broad, ovate, only last whorl visible (SL/MD = 1.5– 1.6). Last whorl globose, periphery placed just above mid-whorl (SL/ PH = 1.8). Apex rounded, deeply and narrowly umbilicate, moderately narrow (MD/ApD = 2.7–2.8), low angled apical plane (AP = 8–9°). Sculpture of punctate spiral cords at extremities extending a variable distance towards mid-whorl, surface glossy. Aperture entire length of shell, narrow adapically (MD/AW = 9.9–11.4), widening anteriorly (MD/MAW = 2.1–2.2), with high expansion rate (ER = 4.7–5.1). Outer lip extending adapically only slightly above apex, convex below, rounded abapically. Columella short, strongly excavated. Columellar callus forming prominent, thick, sharply delimited rim, adherent to base. Columellar callus passing into prominent expanded parietal callus with characteristically (for genus) swollen parietal pad in adapical third. No umbilicus or chink. Discussion. Roxania callifera (Boettger, 1907) is based on a rather slender specimen with prominent spiral sculpture, whereas the herein described specimens are comparatively stout and have reduced sculpture mid-whorl. Both bear a low parietal callus pad, which does not extend far over the base and a strongly excavated columella and adherent columellar rim. Roxania callifera differs from Roxania bomba nov. nom. In being slightly less rounded and having the apical tip of the outer lip less alate. As discussed under that species, it is represented by a single specimen, and it is possible that it represents an extreme form of R. callifera. These two species differ from all other Paratethyan congeners in having a parietal pad developed. Roxania callifera differs from Sabatia isseli Bellardi, 1877, from the Mediterranean Pliocene, in its slenderer outline, less expanding aperture and weaker parietal callus (see Landau et al. 2023: pl. 57). In its relatively weak parietal callus, the Paratethyan species is reminiscent of the extant Sabatia nivea (Watson, 1883) from Vanuatu, which is much larger and more globose (see Valdés 2008: fig. 46). Paleoenvironment. The species is documented from deep water and shallow water assemblages. In the Vienna Basin it is most abundant in the clay of the Baden Formation, suggesting outer neritic water depth (Kranner et al. 2021). The occurrence from Cerová (Slovakia) suggests even upper bathyal environments (Harzhauser et al. 2011) Distribution in Central Paratethys. Badenian (Middle Miocene): Vienna Basin: Baden Vöslau, Baden, Steinebrunn (Austria) (Berger 1953); Făget Basin: Lăpugiu de Sus, Coşteiu de Sus (Romania) (Boettger 1907).

Authors 3

1. Mathias Harzhauser

   Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655

2. Bernard Landau Aachen

   University of Lisbon · Instituto Dom Luiz · Naturalis Biodiversity Center

   Affiliation as printed

   Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494

3. Manuel António E. Malaquias

   University of Bergen · Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655 & Department of Natural History, University Museum of Bergen, University of Bergen, PB 7800, 5020 - Bergen, Norway manuel. malaquias @ uib. no; http: // orcid. org / 0000 - 0002 - 9668 - 945 X & Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655