Pyrunculus latesulcatus

Abstract

Pyrunculus latesulcatus (Boettger, 1902) Figs 21A–F * Coelophysis latesulcata n. sp. — Boettger 1902: 179. Retusa latesulcata (Boettger) — Zilch 1934: 277, pl. 22, fig. 16. Retusa (Acrostemma) latesulcata (Boettger) — Berger 1953: 97, pl. 16, fig. 32. R [etusa]. (A [crostemma). latesulcata (Bttg.) — Sieber 1958: 163. Retusa latesulcata (Boettger) 1901 — Bohn-Havas 1973: 1078. non Atys latesulcata (Boettger, 1902) — Landau et al. 2013: 328, pl. 53, fig. 9 [= Acrostemma anatolica nov. sp]. Type material. Lectotype (designated by Zilch 1934: 277), SMF 363159 - 12.2552a (glass plate 397), SL: 4.1 mm, MD: 1.8 mm, Coşteiu de Sus (Romania), illustrated in Zilch (1934: pl. 22, fig. 16), Fig. 21C. The specimen was destroyed during WWII. Illustrated material. NHMW 1855/0045/0258, SL: 4.2 mm, MD: 1.9 mm, Steinebrunn (Austria), Figs 21A 1 – A 3. NHMW 1855/0045/0258a, SL: 4.6 mm, MD: 2.1 mm, Steinebrunn (Austria), (illustrated in Berger 1953: pl. 16, fig. 32), Figs 21B 1 –B 3. NHMW-ZOO-MO-75000-E-47032, SL: 2.4 mm, MD: 1.1 mm, Lăpugiu de Sus (Romania), Figs 21D 1 –D 3. NHMW 2010/0004/1655, SL: 5.2 mm, MD: 2.2 mm, Bad Vöslau (Austria), Figs 21E 1 –E 2. NHMW 1855/0045/0258a, SL: 3.9 mm, MD: 1.8 mm, Steinebrunn (Austria), Figs 21F 1 –F 2. Revised description. Shell small to medium sized, medium-thickness, moderately slender (SL/MD = 2.2–2.3), elongate conical, only last whorl visible. Last whorl with almost straight to faintly concave sides, forming angle of about 16°. Periphery below mid-whorl, placed just above lower third (SL/ PH = 2.7), rounded at moderately contracting base. Apex relatively narrow (MD/ApD = 2.1–2.2), truncated, apical plane slightly oblique (AP = 11– 12°), deeply and narrowly umbilicate, delimited by round edged, faintly swollen, elevated crest. Sculpture of a few, faint, widely spaced spiral grooves, prominent over base, fading towards mid-whorl, subobsolete on adapical third. Growth lines prominent on apical crest.Aperture slightly longer than shell, very narrow down to periphery (MD/AW = 11.8–14.0), abruptly widening in anterior third below periphery (MD/MAW = 2.0–2.1), resulting in high expansion rate (ER = 5.8–6.7). Outer lip thin, extending some distance above apex, narrowly trigonally alate, weakly concave just above periphery. Columella short, with narrow but prominent columellar callus with subobsolete columellar fold. Columellar callus delimiting narrow umbilical chink. Parietal callus not developed. Discussion. Until now, this species was poorly known because it was based on a fragmentary specimen without the outer lip. Later, Berger (1953) gave a drawing of a specimen from Steinebrunn (Austria), which strongly overemphasized spiral sculpture and the strength of the apical crest (see same specimen illustrated herein as Figs 21B 1 –B 3). Pyrunculus latesulcatus (Boettger, 1902) is most like the extant Mediterranean Pyrunculus hoernesi (Weinkauff, 1866). Both have an identical shape and agree in their apertural features. The extant species has a similar variability in spiral sculpture but differs in the much more prominent axial folds or ribs on the apical crest. Paleoenvironment. The occurrence at Steinebrunn (Austria) suggests shallow marine environments (own data M.H.). Distribution in Central Paratethys. Badenian (Middle Miocene): Vienna Basin: Bad Vöslau, Steinebrunn (Austria) (hoc opus); Pannonian Basin: Pécsvárad, Tekeres (Hungary) (Bohn-Havas 1973); Făget Basin: Coşteiu de Sus, Lăpugiu de Sus (Romania) (Zilch 1934).

Authors 3

1. Mathias Harzhauser

   Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655

2. Bernard Landau Aachen

   University of Lisbon · Instituto Dom Luiz · Naturalis Biodiversity Center

   Affiliation as printed

   Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494

3. Manuel António E. Malaquias

   University of Bergen · Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655 & Department of Natural History, University Museum of Bergen, University of Bergen, PB 7800, 5020 - Bergen, Norway manuel. malaquias @ uib. no; http: // orcid. org / 0000 - 0002 - 9668 - 945 X & Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655