Acteocina lichtensteinii

Abstract

Acteocina lichtensteinii (Eichwald, 1830) Figs 35A–D * A [licula]. Lichtensteinii m.— Eichwald 1830: 214. Bulla clandestina Nov.— Dubois de Montpéreux 1831: 49, pl. 1, figs 19–21. Bull [ina] Lichtensteinii m.— Eichwald 1851: 146, pl. 11, figs 19a–b. Bullina Lichtensteinii— Eichwald 1852: pl. 11, figs 19a–b. Bullin [a]. Lichtensteinii m.— Eichwald 1853: 308. Tornatina Lichtensteini [sic] Eichw.— Friedberg 1928: 541, pl. 35, fig. 15. Tornatina Lichtensteini [sic] Eichw.— Friedberg 1938: 160. Acteocina lichtensteini [sic] (Eichwald, 1830)— Zelinskaya et al. 1968: 238, pl. 53, fig. 18. non Bullina clandestina? Dub.— Hauer 1837: 421 [= Acteocina fusiformis (Boettger, 1907)]. non Bullina Lichtensteinii E. — Hauer 1837: 421 [= Acteocina fusiformis (Boettger, 1907)]. Type material. Syntype illustrated in Eichwald (1852: pl. 11, fig. 19), probably Żukowce (Zhukivtsi) (Ukraine), stored in the St. Petersburg State University, Paleontological Museum (Russia). Illustrated material. NHMW-ZOO-MO-75000-E-47062, SL: 3.4 mm, MD: 2.0 mm, Żukowce (Zhukivtsi) (Ukraine), Figs 35A 1 –A 3. NHMW 1859/9945/0507, SL: 4.0 mm, MD: 2.4 mm, Golubytsya (Hołubica), (Ukraine), Figs 35B 1 –B 2. NHMW 1843/0009/0043, SL: 3.4 mm, MD: 1.9 mm, Żukowce (Zhukivtsi) (Ukraine), Figs 35C 1 –C 4. NHMW-ZOO-MO-75000-E-47067, SL: 4.3 mm, MD: 2.5 mm, Golubytsya (Hołubica) (Ukraine), Figs 35D 1 –D 3. Revised description. Shell small, solid, broad, subcylindrical ovate with slightly convex sides (SL/MD = 1.6–1.8). Protoconch mammillate of about 1.5 whorls, flattened, strongly tilted (~140°), with nucleus more than half covered by first teleoconch whorl. Teleoconch of about 3.5 whorls separated by deeply canaliculated suture. Spire conical (apical angle 89–105°), broad (MD/ApD = 1.5–1.6). Early teleoconch whorls cylindrical with narrow subsutural shelf, rapidly decreasing in height. Periphery placed just above mid-whorl (SL/ PH = 1.7–1.8). Last whorl inflated, 83–90% of total height, rounded shoulder, weakly convex below, rounded at rapidly contracting base. Surface smooth except for delicate, shallowly arcuate growth lines. Aperture 78–83% of total height, moderately narrow in upper third (MD/AW = 7.2–9.3), widening in anterior third (MD/MAW = 2.4–2.8), resulting in moderate expansion rate (ER = 3.0–3.6). Outer lip simple, thin, shallowly arcuate in profile, straight in upper two-thirds, rounded at base. Columella short, moderately excavated, thickened, forming narrow, adherent rim, without or with only subobsolete columellar fold, persisting continuously into narrow parietal callus. Paratethyan synonyms. Bulla clandestina Dubois de Montpéreux, 1831, from Żukowce (Zhukivtsi) (Ukraine), is a subjective junior synonym of Acteocina lichtensteinii (Eichwald, 1830). Discussion. Acteocina lichtensteinii (Eichwald, 1830) differs from all its Paratethyan congeners in its broadly barrel-shaped last whorl. It is known so far only from the late Badenian (Serravallian) of the Polish-Ukrainian Fore-Carpathian Basin, which was a hotspot of endemism at that time (Harzhauser et al. 2024a). Acteocina lichtensteinii is most probably another of these endemic species, which did not spread to other Paratethyan basins. Paleoenvironment. Inner neritic, shallow marine based on associated fauna (own data M.H.). Distribution in Central Paratethys. Badenian (Middle Miocene): Voronyaky Hills: Golubytsya (Hołubica), Yaseniv (Ukraine); Polish-Ukrainian Fore-Carpathian Basin: Biłka, Oles’ko, Tarnoruda, Zalistsi, Zhukivtsi, ravine Zhabyak (Ukraine) (Friedberg 1938).

Authors 3

1. Mathias Harzhauser

   Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655

2. Bernard Landau Aachen

   University of Lisbon · Instituto Dom Luiz · Naturalis Biodiversity Center

   Affiliation as printed

   Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494

3. Manuel António E. Malaquias

   University of Bergen · Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655 & Department of Natural History, University Museum of Bergen, University of Bergen, PB 7800, 5020 - Bergen, Norway manuel. malaquias @ uib. no; http: // orcid. org / 0000 - 0002 - 9668 - 945 X & Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655