Scaphander helveticus Peyrot 1932

Abstract

Scaphander helveticus Peyrot, 1932 Figs 56D 1 –D 2 * Scaphander lignarius Linné mut. helvetica nov. mut.— Peyrot 1932: 204, nr. 1411, pl. 12, figs 25–26. Scaphander lignarius Linné, 1758 —Harzhauser 2002: 127, pl. 11, fig. 17 [non Scaphander lignarius (Linnaeus, 1758)]. Type material. Lectotype (designated herein), specimen illustrated in Peyrot (1932: pl. 12, figs 25–26), Serravallian (Middle Miocene), Salles (France). Illustrated material. NHMW 2000z0001/0053, SL: 46.0 mm, MD: 28.0 mm, Niederkreuzstetten (Austria) (illustrated in Harzhauser 2002: pl. 11, fig. 17), Figs 56D 1 –D 2. Revised description. Shell very large, solid, broad (SL/MD = 1.6), narrow posteriorly, strongly expanding abapically, slightly antero-dorsally compressed. Only last whorl visible. Apex narrow (MD/ApD = 4.6), slightly truncated, apical plane strongly oblique (AP = 18°), apical umbilicus fully sealed, delimited by cord-like, narrowly rounded crest. Periphery placed mid-whorl (SL/ PH = 2.2), weakly concave sides below apex, forming angle of about 55°. Rounded at strongly flaring base. Sculpture of close-set, relatively regular, indistinctly punctate, spiral grooves, delimiting broad, flat spiral cords. Spiral grooves occasionally somewhat irregular at intersections with growth lines. Punctuations not discernible as individual pits. Spiral cords even closer-set at apex. Aperture slightly longer than shell, moderately wide in adapical third (MD/AW = 5.8), widening greatly anteriorly (MD/MAW = 1.4), resulting in high expansion rate (ER = 4.1). Outer lip slightly alate and pointed apically, expanding and almost straight below, broadly rounded abapically. Basal lip solid, Columella deeply and broadly excavated. Columellar callus forming thick, broad, sharply delimited, adherent rim continuing into parietal callus of similar width and strength. Discussion. Scaphander helveticus Peyrot, 1932 differs from Scaphander grundincola (De Gregorio, 1885) in its very broad, drop-shaped outline with faintly concave sides below the apex, more crowded spiral cords, and the thickened apical margin bordering the apical umbilicus. The type specimen from Salles (France) differs from the Paratethyan shell only in its slightly broader spiral cords. Specimens from the Atlantic Upper Miocene of Cacela illustrated by Dollfus et al. (1903: 22, pl. 36, figs 13–14) as Scaphander lignarius were discussed by Landau et al. (2013: 335). They are not that species, neither are they conspecific with any of the species discussed herein. They probably represent an undescribed species that will be discussed in a monograph describing the gastropod assemblage of the locality (Landau et al., in prep). Paleoenvironment. Shallow marine, lagoonal (Harzhauser et al. 2002). Distribution in Central Paratethys. Karpatian (Early Miocene): Korneuburg Basin: Niederkreuzstetten (Austria) (hoc opus). Northeastern Atlantic. Serravallian (Middle Miocene): Aquitaine Basin: Salles, Sallespisse, Manciet (France) (Peyrot 1932).

Authors 3

1. Mathias Harzhauser

   Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655

2. Bernard Landau Aachen

   University of Lisbon · Instituto Dom Luiz · Naturalis Biodiversity Center

   Affiliation as printed

   Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494

3. Manuel António E. Malaquias

   University of Bergen · Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655 & Department of Natural History, University Museum of Bergen, University of Bergen, PB 7800, 5020 - Bergen, Norway manuel. malaquias @ uib. no; http: // orcid. org / 0000 - 0002 - 9668 - 945 X & Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655