Pyrunculus leuchsi

Abstract

Pyrunculus leuchsi (Berger, 1949) Figs 22A–E * Retusa (Cylichnina) leuchsi n. sp. — Berger 1949: 83 (pars). Retusa (Cylichnina) leuchsi n. sp. — Berger 1953: 98 (pars), pl. 17, figs 38–39. R [etusa]. (C [ylichnina]). leuchsi Berg.— Sieber 1958: 163 (pars). Pyrunculus elongatus (Eichwald, 1850) — Švagrovský 1984: 192, pl. 3, fig. 7 [non Pyrunculus elongatus (Eichwald, 1830)]. non Retusa (Cylichnina) elongata leuchsi Berger, 1953 — Strausz 1966: 475, text-fig. 212c [= Retusa sp.]. Type material. Holotype, NHMW-ZOO-MO-75000-E-11815A, SL: 3.0 mm, MD: 1.5 mm, Bad Vöslau (Austria), Figs 22A 1 –A 3. Paratypes, NHMW-ZOO-MO-75000-E-11815, SL: 2.9 mm, MD: 1.4 mm, Bad Vöslau (Austria), Figs 22B 1 –B 3. NHMW-ZOO-MO-75000-E-11815, SL: 3.5 mm, MD: 1.7 mm, Bad Vöslau (Austria), Figs 22C 1 –C 3. Illustrated material. NHMW-ZOO-MO-75000-E-3308, SL: 4.1 mm, MD: 2.0 mm, Bad Vöslau (Austria), Figs 22D 1 –D 3. NHMW-ZOO-MO-75000-E-3308, SL: 4.3 mm, MD: 1.9 mm, Bad Vöslau (Austria), Figs 22E 1 –E 3. Revised description. Shell small, glossy, moderately slender (SL/MD = ~2.2), moderately pyriform, only last whorl visible. Last whorl with short conical apical region (47–50°) passing into almost straight to weakly convex sides, forming angle of about 18–20°, with periphery below mid-whorl (SL/ PH = 2.7–2.8), rounded at moderately contracting base. Apex narrow (MD/ApD = 2.7–3.0), truncated, apical plane slightly oblique (AP = 10–12°), deeply and narrowly umbilicate, delimited by round-edged, elevated crest. Sculpture of a few, shallow spiral grooves at both extremities, rarely persisting to mid-whorl. Aperture longer than shell, very narrow just below mid-whorl (MD/AW = 8.0–10.0), widening anteriorly (MD/MAW = 1.8–2.0), resulting in high expansion rate (ER = 4.0–5.4). Outer lip thin, apically extending some distance above apex, faintly pinched inwards above periphery. Columella short, with narrow columellar callus. Columellar fold, low, weak or subobsolete. Columellar callus adherent or erect, delimiting narrow umbilical chink. Parietal callus not developed. Discussion. Pyrunculus leuchsi (Berger, 1949) is strongly reminiscent of Pyrunculus szobiensis sp. nov. concerning the changing angle between apical region and sides, but it is slenderer, the apex is narrower, and the mid-portion is less cylindrical. Pyrunculus parangistoma (Boettger, 1902) is even more conical and has a narrower apex and Pyrunculus elongatus (Eichwald, 1830) differs in its much higher and slenderer shell. Paleoenvironment. The occurrence in the Baden Formation at Bad Vöslau (Austria) suggests middle to outer neritic environments with up to 210 m water depth (Kranner et al. 2021). Distribution in Central Paratethys. Badenian (Middle Miocene): Vienna Basin: Bad Vöslau (Austria) (Berger 1953), Borský Mikuláš (Slovakia) (Švagrovský 1984).

Authors 3

1. Mathias Harzhauser

   Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655

2. Bernard Landau Aachen

   University of Lisbon · Instituto Dom Luiz · Naturalis Biodiversity Center

   Affiliation as printed

   Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494

3. Manuel António E. Malaquias

   University of Bergen · Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655 & Department of Natural History, University Museum of Bergen, University of Bergen, PB 7800, 5020 - Bergen, Norway manuel. malaquias @ uib. no; http: // orcid. org / 0000 - 0002 - 9668 - 945 X & Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655