Ringiculocosta cancellarioides

Abstract

Ringiculocosta cancellarioides (Seguenza, 1881) Figs 12C, E, 13A Ringicula cancellarioides n. sp. 1879— Seguenza 1881: 44, pl. 2, figs 11, 11a. Ringicula (Ringiculocosta) costata cancellarioides Seguenza— Berger 1954: 134, pl. 12, figs 68–70. R [ingicula]. (R [ingiculocosta.) costata cancellarioides Seg.— Sieber 1958: 163. Ringicula (Ringiculocosta) costata Eichwald, (1830) 1853— Strausz 1966: 471, pl. 36, figs 13–13 [non Ringiculocosta costata (Eichwald, 1830)]. Ringicula costata Eichwald 1830 — Krach 1981: 79, pl. 23, fig. 12 [non Ringiculocosta costata (Eichwald, 1830)]. non Ringicula costata cancellarioides Seguenza— Kókay 1996: 458, pl. 4, fig. 7 [= Ringiculocosta costata (Eichwald, 1830)]. Type specimen. Holotype, SL: 3 mm, MD: 1.7 mm, Sassuolo (Italy), Tortonian (Late Miocene), illustrated in Seguenza (1881: pl. 2, figs 11, 11a). We are not aware of the whereabouts of the type material. Illustrated material. NHMW-ZOO-MO-75000-E-3283, SL: 3.7 mm, MD: 1.8 mm, Bad Vöslau (Austria), illustrated in Berger (1954, pl. 12, fig. 69), Figs 12C 1 –C 2, 13A. NHMW-ZOO-MO-75000-E-57959, SL: 2.6 mm, MD: 1.5 mm, Grund (Austria), Figs 12E 1 –E 2. Revised description. Shell small, moderately broad fusiform (SL/MD: 1.8–1.9) with moderately high conical spire (AA = 38–39°). Protoconch paucispiral of about 1.5 whorls (dp = 400 μm). Teleoconch of three whorls separated by narrowly impressed, linear suture. Spire whorls high, moderately convex with rounded shoulder. Sculpture of close-set axial ribs separated by slightly wider interspaces, overrun by weaker spiral cords, resulting in cancellate pattern. Last whorl 65–66% of total height, with periphery at or just below mid-whorl (SL/ PH = 2.1–2.4), faintly concave subsutural ramp, rounded shoulder, moderately convex periphery; axial interspaces widening on last whorl, spiral cords broaden with narrower interspaces, resulting in rectangular pattern; base conical, constricted. Aperture 49–50% of total height, moderately narrow, widening abapically (MD/MAW = 3.4); insertion of outer lip distinctly below shoulder, thickened by convex labial varix; anal sinus broad, shallow, U-shaped; siphonal canal very short, open, broad, notched at tip. Columella with two strongly elevated, robust folds abapically, parietal denticle prominent, trigonal. Columellar and parietal callus thickened, sharply delimited, narrowly expanded over base. Discussion. This species differs from Ringiculocosta costata (Eichwald, 1830) in its slenderer shell, and the much lower point of insertion of the outer lip. In addition, the two species seem to have been ecologically separated with Ringiculocosta costata in shallow water environments and Ringiculocosta cancellarioides in offshore habitats. Most specimens have a prominent, rectangular sculpture (Fig. 12C) but specimens with weak sculpture also occur (Fig. 12E). Ringiculocosta plicatula (Mayer, 1871), from the Burdigalian of France, differs only in its more prominent axial sculpture (see Mayer 1871: 347, pl. 9, fig. 8; Morlet 1878: 267, pl. 6, fig. 9; Peyrot 1932: 148, pl. 11, figs 29–31). It is possible that R. cancellarioides will become a subjective junior synonym of R. plicatula after revision of French and Italian specimens. Paleoenvironment. The occurrence in the Baden Formation at Bad Vöslau (Austria) suggests middle to outer neritic environments with up to 210 m water depth (Kranner et al. 2021). Distribution in Central Paratethys. Badenian (Middle Miocene): Roztocze Hills: Węglinek, (Poland) (Krach 1981); North Alpine-Carpathian Foreland Basin: Grund (Austria) (hoc opus); Vienna Basin: Bad Vöslau (Austria) (hoc opus); Pannonian Basin: Várpalota (Hungary) (Strausz 1966). Proto-Mediterranean Sea. Tortonian (Late Miocene): Emilia-Romagna: Sassuolo (Italy) (Seguenza 1881).

Authors 3

1. Mathias Harzhauser

   Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655

2. Bernard Landau Aachen

   University of Lisbon · Instituto Dom Luiz · Naturalis Biodiversity Center

   Affiliation as printed

   Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494

3. Manuel António E. Malaquias

   University of Bergen · Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655 & Department of Natural History, University Museum of Bergen, University of Bergen, PB 7800, 5020 - Bergen, Norway manuel. malaquias @ uib. no; http: // orcid. org / 0000 - 0002 - 9668 - 945 X & Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655