Retusa vindobonensis

Abstract

Retusa vindobonensis (Berger, 1949) Figs 31A–C * Retusa (Retusa) vindobonensis n. sp. — Berger 1949: 82 (pars). Retusa (Retusa) vindobonensis n. sp. — Berger 1953: 92 (pars, holotype only), pl. 16, fig. 22, text-fig. 4. Retusa (Cylichnina) pliosimplex (Sacco) — Berger 1953: 99, pl. 17, figs 45–46 [non Retusa pliosimplex (Sacco, 1897)]. R [etusa]. (R [etusa].) vindobonensis Berg.— Sieber 1958: 163. R [etusa]. (C [ylichnina].) pliosimplex (Sacco) — Sieber 1958: 163 [non Retusa pliosimplex (Sacco, 1897)]. Retusa vindobonensis Berger— Kókay 1996: 458, pl. 5, fig. 1. Type material. Holotype, NHMW-ZOO-MO-75000-E-3311A, SL: 3.6 mm, MD: 1.7 mm, Bad Vöslau (Austria), illustrated in Berger (1953: pl. 16, fig. 22), Figs 31A 1 –A 3. Berger (1953) mentioned further occurrences from Lăpugiu de Sus (Romania) and Guntersdorf (Austria). After revision we exclude these specimens because they are not conspecific with Retusa vindobonensis. Illustrated material. NHMW-ZOO-MO-75000-E-7041, SL: 3.5 mm, MD: 3.1 mm, Grund (Austria), Figs 31B 1 –B 3. NHMW-ZOO-MO-75000-E-7041, SL: 4.5 mm, MD: 2.0 mm, Grund (Austria), Figs 31C 1 –C 3. Revised description. Shell small, solid, moderately slender (SL/MD = 2.1–2.3). Only last whorl visible. Protoconch unknown. Periphery just below mid-whorl (SL/ PH = 2.2–2.6) with almost straight sides, weakly widening abapically; adapically forming angle of about 10–20°. Rounded at slowly contracting base.Apex truncated, moderately narrow (MD/ApD = 2.1–2.3), apical plane faintly oblique (AP = 7–10°), deeply and narrowly umbilicate, delimited by rounded crest with delicate, fold-like axial ribs on umbilical side. Sculpture of arcuate growth lines; most prominent on apical crest and extremities, subobsolete mid-whorl. Aperture only slightly longer than shell, narrow down to mid-whorl (MD/AW = 9.0–9.5), distinctly widening anteriorly (MD/MAW = 1.8–2.0), resulting in high expansion rate (ER = 3.3–4.7). Outer lip thin, moderately extending above apex. Columella short, nearly straight; columellar fold weak to subobsolete. Columellar callus forming very narrow, adherent rim delimiting faint umbilical chink. Parietal callus not developed. Discussion. This rare species might be related to Retusa sarmatica Berger, 1953 due to its nearly identical apical morphology and prominent axial sculpture. It differs from R. sarmatica in its slenderer shape, more conical last whorl and narrower apex. Retusa subangystoma (d’Orbigny, 1852), from the Early to Late Miocene of France, differs in its more subcylindrical outline and less convex periphery (see Peyrot 1932: pl. 13, figs 34–35; Landau et al. 2020: pl. 15). Berger (1953) confused this species with Retusa pliosimplex (Sacco, 1897) from the Pliocene of Italy. The Italian species has a similar shape but is slightly more elongate and differs in its wider, cylindrical apical umbilicus. In addition, it has a wider abapical aperture (see Sacco 1897: 51, pl. 4, figs 15–18; Ferrero Mortara et al. 1984: pl. 52, figs 10a–c; Chirli 2023: pl. 12, figs 1–6). Paleoenvironment. The occurrence in the Baden Formation at Bad Vöslau (Austria) suggests middle to outer neritic environments with up to 210 m water depth (Kranner et al. 2021). Distribution in Central Paratethys. Badenian (Middle Miocene): North Alpine-Carpathian Foreland Basin: Grund, Guntersdorf (Austria) (Berger 1953); Vienna Basin: Bad Vöslau (Austria) (Berger 1953); Pannonian Basin: Budapest (Hungary) (Kókay 1996).

Authors 3

1. Mathias Harzhauser

   Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655

2. Bernard Landau Aachen

   University of Lisbon · Instituto Dom Luiz · Naturalis Biodiversity Center

   Affiliation as printed

   Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494

3. Manuel António E. Malaquias

   University of Bergen · Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655 & Department of Natural History, University Museum of Bergen, University of Bergen, PB 7800, 5020 - Bergen, Norway manuel. malaquias @ uib. no; http: // orcid. org / 0000 - 0002 - 9668 - 945 X & Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655