Cylichna intermedia

Abstract

Cylichna intermedia (Sacco, 1897) Figs 43A–E Bulla cylindroides — Naumann 1852: pl. 49, fig. 4 [non Cylichna cylindroides (Deshayes, 1824)]. [Bullinella] Cylichnina intermedia Sacc.— Sacco 1897: 52, pl. 4, figs 31–32. Retusa (Cylichnina) intermedia (Sacco) — Berger 1953: 101, pl. 19, figs 93–94. R [etusa]. (C [ylichnina].) intermedia (Sacco) — Sieber 1958: 163. Retusa (Cylichnina) intermedia Sacco— Kókay 1996: 458, pl. 5, fig. 9. Cylichnina intermedia Sacco, 1897 — Ferrero Mortara et al. 1984: 288, pl. 53, fig. 1. Bulla (Cylichnina) hungarica (Boettger, 1906) — Atanacković 1985: 189, pl. 42, figs 11–12 [non Bulla hungarica Boettger, 1907]. Cylichna clathrata (Defrance, 1825) — Landau et al. 2013: 331, pl. 52, fig. 18 [non Cylichna clathrata (Defrance, 1814)]. Type material. Syntype, BS.090.02.019, SL: 8.8 mm, MD: 3.8 mm, Tetti Borelli (Italy), Late Miocene, illustrated in Ferrero Mortara et al. (1984: pl. 53, fig. 1). Illustrated material. NHMW-ZOO-MO-75000-E-7083, SL: 8.3 mm, MD: 3.6 mm, Gainfarn (Austria), Figs 43A 1 –A 3. NHMW-ZOO-MO-75000-E-47087, SL: 7.9 mm, MD: 3.7 mm, Steinebrunn (Austria), Figs 43B 1 –B 3. NHMW-ZOO-MO-75000-E-33312, SL: 8.5 mm, MD: 3.9 mm, Guntersdorf (Austria), Figs 43C 1 –C 2. NHMW 1854/0035/0386, SL: 6.7 mm, MD: 3.1 mm, Lăpugiu de Sus (Romania), Figs 43D 1 –D 3. NHMW 1997z0178/1884, SL: 9.2 mm, MD: 4.2 mm, Gainfarn (Austria), Figs 43E 1 –E 3. Revised description. Shell medium sized, solid, moderately slender subcylindrical (SL/MD = 2.2–2.3), only last whorl visible. Last whorl elongate, with weakly convex sides, slowly contracting at base; periphery just above mid-whorl (SL/ PH = 1.8–1.9); apex low, convex, narrow (MD/ApD = 2.8–2.9), weakly oblique (AP = 9–11°), deeply and narrowly umbilicate, delimited by convex ridge. Surface smooth except for fine spirals over base. Aperture as long as the shell, moderately narrow adapically and in mid-section (MD/AW = 6.7–7.0), widening anteriorly (MD/ MAW = 2.2–2.3), with low to moderate expansion rate (ER = 2.9–3.3). Outer lip slightly extending above apex, straight to very weakly convex below, rounded abapically. Columella short, slightly excavated; columellar callus forming relatively broad rim, slightly extending over base, with distinct, narrow columellar fold. Columellar callus short, slightly expanded over medial side of base. No parietal callus. Narrow umbilical chink. Discussion. Cylichna clathrata (Defrance, 1825), from the Early Miocene of France as illustrated by Peyrot (1932: pl. 13, figs 11, 15), differs from Cylichna intermedia (Sacco, 1897) in its nearly straight sided last whorl, has its periphery distinctly below mid-whorl (SL/ PH = 2.5), lacks a columellar fold and has a much wider maximum aperture width (MD/MAW = 1.8) (versus 2.2–2.3 in Cylichna intermedia). Landau et al. (2013: 331, pl. 52, fig. 18) mistakenly attributed specimens from the Serravallian Karaman Basin of Turkey to C. clathrata. The illustrated specimen clearly shows a terminal columellar ridge and they are herein reassigned to C. intermedia. Bulla clathrata sensu de Basterot (1825: pl. 1, fig. 10) has a comparable shape to C. intermedia but lacks a columellar fold. Paleoenvironment. Shallow marine, inner neritic based on the associated fauna (own data, M.H.). Distribution in Central Paratethys. Badenian (Middle Miocene): North Alpine-Carpathian Foreland Basin: Grund, Guntersdorf (Austria) (Berger 1953); Vienna Basin: Mikulov (Czech Republic), Gainfarn, Steinebrunn, Vienna /Pötzleinsdorf (Austria) (Berger 1953); Pannonian Basin: Budapest (Hungary) (Kókay 1996); Southern Pannonian Basin: Hrvaćani, Miljevići (Bosnia and Herzegovina) (Atanacković 1985); Făget Basin: Buituri, Lăpugiu de Sus (Romania) (Berger 1953). Proto-Mediterranean Sea. Serravallian (Middle Miocene): Karaman Basin: Akpınar, Seyithasan (Turkey) (Landau et al. 2013). Tortonian/Messinian (Late Miocene): Colli Torinesi: Tetti Borelli (Italy) (Ferrero Mortara et al. 1984).

Authors 3

1. Mathias Harzhauser

   Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655

2. Bernard Landau Aachen

   University of Lisbon · Instituto Dom Luiz · Naturalis Biodiversity Center

   Affiliation as printed

   Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494

3. Manuel António E. Malaquias

   University of Bergen · Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655 & Department of Natural History, University Museum of Bergen, University of Bergen, PB 7800, 5020 - Bergen, Norway manuel. malaquias @ uib. no; http: // orcid. org / 0000 - 0002 - 9668 - 945 X & Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655