Cosmophiline Berger 1949

Abstract

Genus Cosmophiline Berger 1949 Type species. Cosmophiline marinellii Berger, 1949; by monotypy. Middle Miocene, Austria. Diagnosis. “Shell very thin, small apex, involute, last whorl strongly expanding (as in Philine Ascanius). Surface with strong spiral cords, which diverge arcuately on the last whorl diverge towards the upper and lower margin and terminate in strong teeth.” (Berger 1953: 115; translated from German). Discussion. Oskars et al. (2015) showed the family Philinidae to be polyphyletic with four lineages warranting familial status (Philinidae, Philinorbidae, Laonidae, and an unnamed clade informally designated by the authors as Philinidae Clade 4). Since then, a new family has been added, namely Antarctophilinidae for Southern Ocean species (Moles et al. 2019). Although the separation between these families relies mostly on anatomical characters such as presence/absence of gizzard plates as well as the nature of these plates, shells are also fairly informative for family and generic assignments. The genus of Cosmophiline is characterized by a very distinct shell which to our best knowledge is not present among extant philinid snails. Based on the quadrangular and flattish shape of these shells we assign the genus to the family Philinidae.

Authors 3

1. Mathias Harzhauser

   Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655

2. Bernard Landau Aachen

   University of Lisbon · Instituto Dom Luiz · Naturalis Biodiversity Center

   Affiliation as printed

   Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494

3. Manuel António E. Malaquias

   University of Bergen · Natural History Museum Vienna

   Affiliation as printed

   Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655 & Department of Natural History, University Museum of Bergen, University of Bergen, PB 7800, 5020 - Bergen, Norway manuel. malaquias @ uib. no; http: // orcid. org / 0000 - 0002 - 9668 - 945 X & Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655