Copera vittata subsp. vittata

Zenodo (CERN European Organization for Nuclear Research)

Abstract

Copera vittata vittata (Selys, 1863) Selected references: Copera vittata vittata Selys, 1863;— Lieftinck 1940a: 295 (first record from Riau). Psilocnemis acutimargo Krüger, 1898;— Schmidt 1934: 336 (first record from Sumatra). Specimens collected. M1b: 1 ♂ (RER20 _ PCD5; UMRI), 18 i 2020; 3 ♀♀ (RER20 _ PCD6–8; RER), 18 i 2020; 1 ♂, 1 ♀ (RER20 _ PCD9–10; BRIN), in tandem, 22 i 2020; M1d: 1 ♀ (RER20 _ PCD1; RER), 15 i 2020; M1e: 1 ♂ (RER20 _ PCD11; UMRI), 26 i 2020; M2a: 3 ♂♂ (RER20 _ PCD2–4; BRIN), 16 i 2020; 1 ♂ (RER23 _ PCD93; UMRI), 2 x 2023; M4a: 1 ♂, 1 ♀ (RER23 _ PCD8–9; BRIN), in tandem, 13 ii 2023; 2 ♂♂ (RER23 _ PCD54–55; UMRI), 5 vi 2023; 1 ♂, 1 ♀ (RER23 _ PCD63–64; RER), 14 vi 2023; 1 ♂, 1 ♀ (RER23 _ PCD89–90; UMRI), tandem pair, 27 ix 2023; 1 ♂ (RER23 _ PCD92; BRIN), 27 ix 2023; M4b: 2 ♂♂ (RER23 _ PCD2–3; BRIN), 8 ii 2023; 2 ♂♂ (RER23 _ PCD10–11; UMRI), 13 ii 2023; 1 ♀ (RER23 _ PCD12; RER), teneral, 13 ii 2023, coll. RER staff; 1 ♂, 1 ♀ (RER23 _ PCD58–59; BRIN), tandem pair, 13 vi 2023; 3 ♂♂ (RER23 _ PCD60–62; UMRI), 13 vi 2023; M4e: 1 ♀ (RER23 _ PCD7; RER), 13 ii 2023; 1 ♂ (RER23 _ PCD91; RER), 27 ix 2023; M5b: 2 ♂♂, 1 ♀ (RER23 _ PCD4–6; UMRI), 9 ii 2023; M5f: 1 ♂, 1 ♀ (RER23 _ PCD56–57; RER), tandem pair, 6 vi 2023; M10a: 1 ♂ (RER23 _ PCD1; RER), 7 ii 2023; M12d: 1 ♂ (RER23 _ PCD65; BRIN), 18 vi 2023; P1b: 1 ♂. 1 ♀ (RER23 _ PCD52–53; BRIN), 2 vi 2023; P6f: 1 ♂ (RER23 _ PCD88; RER), 25 ix 2023. Taxonomic remarks. The variety of color forms (and general variability) of C. vittata has been commented on before (for example see Dow & Singa (2022) on forms found in Sarawak and Brunei) but no detailed study of C. vittata and its closest allies has been made since that of Lieftinck (1940a). Two color forms were found at the Kampar Peninsula: (a) a largely dark form with yellow legs and a very variable (sometimes very reduced) pattern of pale markings on the synthorax, this form is very common in the low pH habitats of the main part of the peninsula; (b) the ‘typical’ red-legged form, only a single individual of this form was collected, at a small stream in oil palm on mineral soils at the base of the peninsula. In Borneo typically a single form is found in a given area, for instance a yellow-legged form is found in the southwest of Sarawak (where it occurs in a variety of habitats including PSF), with an exception in Brunei and Limbang Division in northeast Sarawak where the ‘typical’ red-legged form and a black form (Orr 2003, Dow & Singa 2022) both occur, but without clear partitioning by habitat. The situation in the Kampar Peninsula contrasts with that in Borneo with the two forms occurring in the Kampar Peninsula seemingly occupying different habitats. The yellow-legged form (which might be smaller, at least on average, than the ‘typical’ form but detailed measurements are needed to confirm this) present in the peninsula might be a low pH phenotype but the possibility that it is a distinct taxon cannot be excluded and further work is needed on this entire complex.

Authors 7

1. Rory A. Dow Aachen Institute of Biodiversity and Environmental Conservation

   Naturalis Biodiversity Center · Universiti Malaysia Sarawak

   Affiliation as printed

   Institute of Biodiversity and Environmental Conservation, Universiti Malaysia Sarawak, 94300 Kota Samarahan, Sarawak, Malaysia. & Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, The Netherlands

2. Novia Gesriantuti

   Universitas Muhammadiyah Riau

   Affiliation as printed

   Program Studi Biologi Fakultas MIPA Dan Kesehatan, Universitas Muhammadiyah Riau, Indonesia

3. Muhammad Iqbal

   Riau University

   Affiliation as printed

   Restorasi Ekosistem Riau, Pangkalan Kerinci

4. Pungki Lupiyan-Ingdyah

   Brigham Young University · National Research and Innovation Agency

   Affiliation as printed

   Research Center for Biosystematics and Evolution, National Research and Innovation Agency (BRIN), Cibinong, Kabupaten Bogor, Jawa Barat 16911, Indonesia. & Department of Biology, Brigham Young University, Provo, UT 84602, USA

5. JOHN F. PERIERA
   Affiliation as printed

   Restorasi Ekosistem Riau, Pangkalan Kerinci, Riau

6. Shinta O. Zulya

   Universitas Muhammadiyah Riau

   Affiliation as printed

   Program Studi Biologi Fakultas MIPA Dan Kesehatan, Universitas Muhammadiyah Riau, Indonesia

7. Bradford M. Sanders
   Affiliation as printed

   Restorasi Ekosistem Riau, Pangkalan Kerinci, Riau